Coleoptera (Beetles)
Family: Anthicidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Anthicodes maculatus | Wollaston, 1877 | Spotted ant-like beetle | 1876 | St. Helena | Missing | Access | 
Family: Anthribidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Homoeodera asteris | Wollaston, 1877 | Flagstaff fungus weevil, Scrubwood fungus beetle | 1876 | St. Helena | Missing or invalid | Access | 
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 Homoeodera nodulipennis  | 
 Wollaston, 1877  | 
 Gnarled broad-headed weevil  | 
 pre-1878  | 
 St. Helena  | 
 Extinct  | 
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 Urodontidius enigmaticus  | 
 Louw, 1993  | 
 -  | 
 (rediscovered)  | 
 South Africa  | 
 Rediscovered  | 
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| Valenfriesia congener | (Wollaston, 1877) | Blackened fungus weevil, Congeneric fungus beetle | March 1967 | St. Helena | Missing | Access | 
| 
 Valenfriesia rotundata  | 
 (Wollaston, 1877)  | 
 Rotund fungus weevil, Rotund fungus beetle  | 
 March 1967  | 
 St. Helena  | 
 Missing  | 
Family: Bothrideridae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| 
 Ascetoderes strigatus  | 
 (Arrow in Waterhouse et al., 1900)  | 
 -  | 
 1897-1898  | 
 Christmas Island, Australia  | 
 Missing  | 
Family: Brachinidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Brachinus solidipalpis | Tian & Deuve, 2007 | - | 1843 | Manila, Philippines | Missing | Access | 
Family: Buprestidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Agrilus toyoshimai | Toyama, 1988 | (rediscovered) | Honshu, Japan | Invalid (synonym) | Access | |
| Castiarina insculpta | Carter, 1934 | Miena jewel beetle | (rediscovered) | Central Plateau, Tasmania, Australia | Rediscovered | Access | 
| Melobasis empyria | Olliff, 1889 | Fiery jewel beetle | ? | Lord Howe Island, New South Wales, Australia | Missing | Access | 
Family: Carabidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| 
 Bembidion derelictus  | 
 Alluaud, 1926  | 
 -  | 
 1888  | 
 Flores, Azores  | 
 Extinct  | 
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 Bembidion dicksoniae  | 
(Wollaston, 1877) | 1875-1876 | St. Helena | Missing | Access | |
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| Bembidion fossor | (Wollaston, 1877) | 1875-1876 | St. Helena | Missing | Access | |
| Bembidion megalops | (Wollaston, 1877) | 1884-1886 | St. Helena | Missing | Access | |
| 
 Bembidion palosverdes  | 
 Kavanaugh & Erwin, 1992  | 
 Palos Verdes intertidal minute carabid beetle  | 
 (rediscovered)  | 
 Palos Verdes Peninsula, Los Angeles County, California, USA  | 
 Rediscovered  | 
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 Bembidion sublimbatum  | 
 (Wollaston,1877)  | 
 -  | 
 1884-1886  | 
 High Central Ridge, St Helena  | 
 Missing  | 
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 Bembidion trechoides  | 
(Wollaston, 1877) | 1875-1876 | St. Helena | Missing | Access | |
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 Blackburnia agonoides  | 
 Sharp, 1903  | 
 -  | 
 1896  | 
 Hawaiian Islands, USA  | 
 Missing  | 
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 Blackburnia anomala  | 
 (Blackburn, 1878)  | 
 -  | 
 (rediscovered)  | 
 Maui, Hawaiian Islands, USA  | 
 Rediscovered  | 
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 Blackburnia blaptoides  | 
 Blackburn, 1878  | 
 -  | 
 pre-1901  | 
 Hawaiian Islands, USA  | 
 Extinct  | 
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 Blackburnia burneyi  | 
 Liebherr & Porch, 2015  | 
 -  | 
 late Holocene  | 
 Kauai, Hawaiian Islands, USA  | 
 Extinct  | 
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 Blackburnia cryptipes  | 
 Liebherr & Porch, 2015  | 
 -  | 
 late Holocene  | 
 Kauai, Hawaiian Islands, USA  | 
 Extinct  | 
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| Blackburnia debilis | (Perkins, 1917) | (rediscovered) | Kauai, Hawaiian Islands, USA | Rediscovered | Access | |
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 Blackburnia godzilla  | 
 Liebherr & Porch, 2015  | 
 -  | 
 late Holocene  | 
 Kauai, Hawaiian Islands, USA  | 
 Extinct  | 
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 Blackburnia haleakala  | 
 Liebherr, 2000  | 
 -  | 
 1896  | 
 Hawaiian Islands, USA  | 
 Extinct  | 
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 Blackburnia insignis  | 
 Sharp, 1878  | 
 -  | 
 ?  | 
 Hawaiian Islands, USA  | 
 Missing  | 
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 Blackburnia lenta  | 
 Sharp, 1903  | 
 -  | 
 (rediscovered)  | 
 Hawaiian Islands, USA  | 
 Rediscovered  | 
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 Blackburnia menehune  | 
 Liebherr & Porch, 2015  | 
 -  | 
 late Holocene  | 
 Kauai, Hawaiian Islands, USA  | 
 Extinct  | 
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 Blackburnia metromenoides  | 
 Perkins, 1917  | 
 Waianae Mountains ground beetle  | 
 ?  | 
 Oahu, Hawaiian Islands, USA  | 
 Extinct  | 
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 Blackburnia micantipennis  | 
 Sharp, 1903  | 
 Kauai ground beetle, Waimea ground beetle  | 
 1935  | 
 Kauai, Hawaiian Islands, USA  | 
 Extinct  | 
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 Blackburnia mothra  | 
 Liebherr & Porch, 2015  | 
 -  | 
 late Holocene  | 
 Kauai, Hawaiian Islands, USA  | 
 Extinct  | 
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 Blackburnia mystica  | 
 Blackburn, 1877  | 
 Mystical ground beetle  | 
 ?  | 
 Hawaiian Islands, USA  | 
 Missing  | 
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 Blackburnia oceanica  | 
 Blackburn, 1877  | 
 Koolau Mountains ground beetle, Oceanic ground beetle  | 
 ?  | 
 Oahu, Hawaiian Islands, USA  | 
 Missing  | 
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 Blackburnia octoocellata  | 
 (Karsch, 1881)  | 
 -  | 
 1902  | 
 Hawaiian Islands, USA  | 
 Missing  | 
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 Blackburnia ovata  | 
 Liebherr & Porch, 2015  | 
 -  | 
 late Holocene  | 
 Kauai, Hawaiian Islands, USA  | 
 Extinct  | 
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 Blackburnia rugosa  | 
 Liebherr & Porch, 2015  | 
 -  | 
 late Holocene  | 
 Kauai, Hawaiian Islands, USA  | 
 Extinct  | 
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 Blackburnia sharpi  | 
 (Blackburn, 1878)  | 
 -  | 
 1902  | 
 Hawaiian Islands, USA  | 
 Missing  | 
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 Blackburnia tantalus  | 
 (Blackburn, 1877)  | 
 Oahu ground beetle  | 
 ?  | 
 Hawaiian Islands, USA  | 
 Missing  | 
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 Blackburnia terebrata  | 
 Blackburn, 1881  | 
 -  | 
 1896  | 
 Hawaiian Islands, USA  | 
 Missing  | 
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 Blackburnia transiens  | 
 (Sharp, 1903)  | 
 Transient ground beetle  | 
 ?  | 
 Kauai, Hawaiian Islands, USA  | 
 Missing  | 
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 Bradycellus chavesi  | 
 Alluaud, 1919  | 
 -  | 
 1918  | 
 São Miguel Island, Azores  | 
 Extinct  | 
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 Bryanites graeffii  | 
 Liebherr, 2017  | 
 -  | 
 pre-1871  | 
 Samoa  | 
 Extinct  | 
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 Calathus extensicollis  | 
 Putzeys, 1873  | 
 -  | 
 1930  | 
 Pico, Azores  | 
 Extinct  | 
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 Calathus vicenteorum  | 
 Schatzmayr, 1937  | 
 -  | 
 1985  | 
 São Miguel Island, Azores  | 
 Extinct  | 
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| Calosoma burchellii | (Waterhouse, 1842) | Giant ground beetle, Burchell's Giant ground beetle | 1967 | St. Helena | Extinct | Access | 
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 Cicindelidia floridana  | 
 Cartwright, 1939  | 
 Miami scabrous tiger beetle, Floridan tiger beetle, Miami tiger beetle  | 
 (rediscovered)  | 
 Miami, Florida, USA  | 
 Rediscovered  | 
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 Cicindela chlorocephala smythi  | 
 Harris, 1913  | 
 Smyth's tiger beetle, Smyth’s lime-headed tiger beetle  | 
 June 1912  | 
 Padre Island, Texas, USA  | 
 Extinct  | 
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 Cicindela hirticollis abrupta  | 
 Casey, 1913  | 
 Sacramento Valley hairy-necked beetle  | 
 c.1984  | 
 Sacramento Valley, California, USA  | 
 Extinct  | 
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 Cicindela latesignata obliviosa  | 
 Casey, 1913  | 
 Oblivious tiger beetle, Western beach tiger beetle  | 
 c.1980?  | 
 California, USA  | 
 Missing or invalid (synonym)  | 
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 Cicindela tranquebarica joaquinensis  | 
 Knisley & Haines, 2007  | 
 San Joaquin tiger beetle  | 
 ?  | 
 California, USA  | 
 Rediscovered  | 
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 Clivina morio  | 
 Dejean, 1831  | 
 -  | 
 (rediscovered)  | 
 USA  | 
 Rediscovered  | 
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 Cylindera uudata  | 
 (Horn, 1915)  | 
 -  | 
 (rediscovered)  | 
 Durango, Mexico  | 
 Rediscovered  | 
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 Cymindis alutacea  | 
 Wollaston, 1867  | 
 -  | 
 (rediscovered)  | 
 São Nicolau, Cape Verde Islands  | 
 Rediscovered  | 
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 Elaphrus viridis  | 
 Horn, 1878  | 
 
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 (rediscovered)  | 
 Solano County, California, USA  | 
 Rediscovered  | 
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 Harpalus prosperus  | 
 Basilewsky, 1972  | 
 -  | 
 1960's  | 
 St. Helena  | 
 Rediscovered  | 
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 Holcaspis brevicula  | 
 Butcher, 1984  | 
 Eyrewell ground beetle  | 
 2005  | 
 Eyrewell Forest, Canterbury Plains, New Zealand  | 
 Missing  | 
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 Horologion speokoites  | 
 Valentine, 1932  | 
 Arbuckle Cave ground beetle  | 
 ?  | 
 West Virginia, USA  | 
 Missing  | 
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 Ishikawatrechus intermedius  | 
 Ueno, 1957  | 
 Kadota Chibi blind ground beetle  | 
 (rediscovered)  | 
 Kôchi Prefecture, Shikoku, Japan  | 
 Rediscovered  | 
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 Lacordairia fugax  | 
 (Olliff, 1889)  | 
 -  | 
 ?  | 
 Lord Howe Island, New South Wales, Australia  | 
 Missing  | 
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 Laemostenus libanensis  | 
 (Piochard de la Brûlerie, 1876)  | 
 -  | 
 (rediscovered)  | 
 Lebanon  | 
 Rediscovered  | 
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 Leistus punctatissimus  | 
 Breit, 1914  | 
 -  | 
 (rediscovered)  | 
 Italy  | 
 Rediscovered  | 
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 Mecodema costellum sp. nov. 'South Canterbury'  | 
 -  | 
 South Canterbury weevil  | 
 ?  | 
 South Canterbury, South Island, New Zealand  | 
 Missing  | 
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 Mecodema costellum ssp. nov. 'spelles'  | 
 -  | 
 Paturau Caves mecodema weevil  | 
 Before 1840?  | 
 Paturau Caves, Nelson (northwest), South Island, New Zealand  | 
 Extinct  | 
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 Mecodema punctellum  | 
 Broun, 1921  | 
 Stephens Island weevil  | 
 1931  | 
 Stephens Island, New Zealand  | 
 Extinct  | 
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 Mecodema sp. nov. 'South Canterbury'  | 
 -  | 
 South Canterbury weevil  | 
 ?  | 
 South Canterbury, South Island, New Zealand  | 
 Missing  | 
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 Mecyclothorax nubicola  | 
 (Blackburn, 1878)  | 
 -  | 
 (rediscovered)  | 
 Hawaiian Islands, USA  | 
 Rediscovered  | 
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 Mecyclothorax rusticus  | 
 Sharp, 1903  | 
 -  | 
 (rediscovered)  | 
 Hawaiian Islands, USA  | 
 Rediscovered  | 
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 Megadromus sp. nov. 'South Island'  | 
 -  | 
 -  | 
 ?  | 
 South Island, New Zealand  | 
 Missing  | 
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 Metromenus cuneipennis  | 
 Blackburn, 1877  | 
 Wedge-winged metromenus ground beetle  | 
 (rediscovered)  | 
 O'ahu, Hawaiian Islands, USA  | 
 Rediscovered  | 
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 Nurus atlas  | 
 (Laporte, 1867)  | 
 -  | 
 (rediscovered)  | 
 Australia  | 
 Rediscovered  | 
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 Nurus brevis  | 
 Motschulsky, 1865  | 
 -  | 
 (rediscovered)  | 
 Australia  | 
 Rediscovered  | 
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 Platynus dissipatus  | 
 Scudder, 1890  | 
 -  | 
 Late Pleistocene  | 
 Canada  | 
 Extinct  | 
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 Pseudanophthalmus fowlerae  | 
 Barr, 1980  | 
 Fowler's cave beetle  | 
 (rediscovered)  | 
 Tennessee, USA  | 
 Rediscovered  | 
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 Pseudanophthalmus insularis  | 
 Barr, 1959  | 
 Bakers Station cave beetle  | 
 (rediscovered)  | 
 Tennessee, USA  | 
 Rediscovered  | 
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 Pseudanophthalmus krameri  | 
 Krekeler, 1973  | 
 Kramer's cave beetle  | 
 ?  | 
 Cave Hill Cave, Ohio, USA  | 
 Extinct  | 
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 Pseudanophthalmus krekeleri  | 
 Barr, 1965  | 
 Rich Mountain cave beetle  | 
 1985  | 
 West Virginia, USA  | 
 Extinct  | 
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 Pseudanophthalmus parvus  | 
 Krekeler, 1973  | 
 Tatum Cave beetle  | 
 1965  | 
 Marion County, Kentucky, USA  | 
 Missing  | 
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 Pseudanophthalmus paulus  | 
 Barr, 1981  | 
 Noblett's cave beetle  | 
 (rediscovered)  | 
 Tennessee, USA  | 
 Rediscovered  | 
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 Pseudanophthalmus tiresias  | 
 Barr, 1959  | 
 Soothsayer cave beetle  | 
 (rediscovered)  | 
 Tennessee, USA  | 
 Rediscovered  | 
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 Pseudanophthalmus tullahoma  | 
 Barr, 1959  | 
 Duck River Cave ground beetle  | 
 ?  | 
 Tennessee, USA  | 
 Missing  | 
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| Pseudomyas doramasensis | Uyttenboogaart, 1929 | 1927 | Gran Canaria, Canary Islands | Missing | Access | |
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 Pterostichus destitutus  | 
 Scudder, 1890  | 
 -  | 
 Late Pleistocene  | 
 Canada  | 
 Extinct  | 
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 Rakantrechus elegans  | 
 Uéno, 1960  | 
 Kozono blind chibi beetle  | 
 1955?  | 
 Kozono, Tsukumi city, Oita Prefecture, Kyushu, Japan  | 
 Missing  | 
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 Rhadine ozarkensis  | 
 Sanderson & Miller, 1941  | 
 Ozark Cave ground beetle  | 
 (rediscovered)  | 
 Fincher('s) Cave, Washington County, Arkansas, USA  | 
 Rediscovered  | 
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 Trechus balsamensis  | 
 Barr, 1962  | 
 Plott Balsam Mountains ground beetle  | 
 ?  | 
 North Carolina, USA  | 
 Extinct  | 
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 Trechus carolinae  | 
 Schaeffer, 1901  | 
 Black Mountains ground beetle  | 
 ?  | 
 North Carolina, USA  | 
 Extinct  | 
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 Trechus luculentus luculentus  | 
 Barr, 1962  | 
 Bright ground beetle  | 
 ?  | 
 North Carolina & Tennessee, USA  | 
 Extinct  | 
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 Trechus luculentus unicoi  | 
 Barr, 1979  | 
 Unicoi Mountains ground beetle  | 
 ?  | 
 North Carolina & Tennessee, USA  | 
 Extinct  | 
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 Trechus mitchellensis  | 
 Barr, 1962  | 
 Mitchell's ground beetle  | 
 ?  | 
 North Carolina, USA  | 
 Extinct  | 
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 Trechus novaculosus  | 
 Barr, 1962  | 
 Great Smoky Mountains ground beetle  | 
 ?  | 
 North Carolina & Tennessee, USA  | 
 Extinct  | 
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 Trechus roanicus  | 
 Barr, 1962  | 
 Roan Mountain ground beetle  | 
 ?  | 
 North Carolina & Tennessee, USA  | 
 Extinct  | 
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 Trechus rosenbergi  | 
 Barr, 1962  | 
 Rosenberg's ground beetle  | 
 ?  | 
 North Carolina, USA  | 
 Extinct  | 
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 Trechus satanicus  | 
 Barr, 1962  | 
 Satanic ground beetle  | 
 ?  | 
 North Carolina, USA  | 
 Extinct  | 
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 Trechus subtilis  | 
 Barr, 1962  | 
 Fine ground beetle  | 
 ?  | 
 North Carolina, USA  | 
 Extinct  | 
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 Trechus torretassoi  | 
 Jeannel, 1937  | 
 -  | 
 1985  | 
 São Miguel Island, Azores  | 
 Extinct  | 
Family: Cerambycidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Anoplophora ogasawaraensis | Makihara, 1976 | Ogasawara longhorn beetle | 2 June 1915 | Ogasawara Islands (=Bonin Islands) (island unknown) | Missing | Access | 
| Diseoblax wrightii | (Waterhouse, 1880) | Gros moutouc du filao | 1950's? | Seychelles | Extinct | Access | 
| Elasmotena insulana | (Olliff, 1890) | - | ? | Lord Howe Island, New South Wales, Australia | Missing | Access | 
| Oricopis insulana | (Olliff, 1890) | - | (rediscovered) | Lord Howe Island, New South Wales, Australia | Rediscovered | Access | 
| Paradeucalion desertarum | (Wollaston, 1854) | ? | Ihlas Desertas, Madeira, Macaronesia | Missing | Access | |
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 Romulus globosus  | 
Knull, 1948 | - | (rediscovered) | Florida, USA | Rediscovered | Access | 
| Somatidia pulchella | Olliff, 1889 | - | ? | Lord Howe Island, New South Wales, Australia | Missing | Access | 
| 
 Xylotoles costatus  | 
Pascoe, 1875 | Pitt Island longhorn beetle | (rediscovered) | Chatham Islands, New Zealand | Rediscovered | Access | 
| Xylotoles wollastoni | (White, 1856) | - | (rediscovered) | Lord Howe Island, New South Wales, Australia | Rediscovered | Access | 
Family: Chrysomelidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Chrysolina fragariae | (Wollaston, 1854) | - | (rediscovered) | Madeira | Rediscovered | Access | 
| Sphenoraia chujoi | Lee, 2014 | Sôzan beetle | 1936 | Sôzan, Taiwan | Extinct | Access | 
Family: Cicindelidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Jansenia laeticolor | (W. Horn, 1904) | - | (rediscovered) | Sri Lanka | Rediscovered | Access | 
| Pometon bolivianus | Huber, 1999 | Bolivian tiger beetle, Bolivian ornate tiger beetle | (rediscovered) | Bolivia | Rediscovered | Access | 
Family: Cleridae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Cormodes darwinii | Pascoe, 1860 | - | (rediscovered) | Lord Howe Island, New South Wales, Australia | Rediscovered | Access | 
Family: Coccinellidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Henosepilachna nativitatis | (Arrow in Waterhouse et al., 1900) | - | pre-1900 | Christmas Island, Australia | Missing | Access | 
| Micraspis flavovittata | Crotch, 1874 | - | (rediscovered) | Victoria, Australia | Rediscovered | Access | 
Family: Curculionidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Acanthinomerus monilicornis | (Wollaston, 1869) | (rediscovered) | St. Helena | Rediscovered | Access | |
| Anagotus sp. nov. 'Fred Cave' | - | Fred Cave anagotus weevil | ? | Waitomo, North Island, New Zealand | Extinct | Access | 
| Anchonocranus oleae | Marshall, 1912 | (rediscovered) | South Africa | Rediscovered | Access | |
| Bagous lyali | Caldara & O'Brien, 1998 | - | 5 July 1934 | Hula Valley, Israel | Missing | Access | 
| Cossonus variipennis | Gahan, 1900 | - | ? | Christmas Island, Australia | Missing | Access | 
| Cryptommata cuccalata | Wollaston, 1877 | ? | St. Helena | Missing | Access | |
| Curculio caryatrypes | (Boheman, 1843) | Larger chesnut weevil | 1956 | Canada & USA | Extinct | Access | 
| Dryophthorus distinguendus | Perkins, 1900 | - | 1961 | Hawaiian Islands, USA | Extinct | Access | 
| Dryotribus mimeticus | Horn, 1873 | - | 1953 | French Frigate Shoals, Johnston Island, Laysan and Wake Island, Hawaiian Islands, USA | Extinct | Access | 
| Dryotribus wilderi | Perkins, 1916 | - | June 1962 | Hawaiian Islands, USA (French Frigate Shoals, Kure, Laysan & Pearl and Hermes) | Missing | Access | 
| Hadramphus tuberculatus | Pascoe, 1877 | Canterbury knobbed weevil, Spaniard weevil, Banks Peninsula speargrass weevil | (rediscovered) | Canterbury Plains, South Island, New Zealand | Rediscovered | Access | 
| Hadramphus sp. nov. 'Westland caves' | - | - | subfossil | Westland Caves area, South Island, New Zealand | Missing | Access | 
| Howeotranes insularis | (Pascoe, 1874) | - | (rediscovered) | Lord Howe Island, New South Wales, Australia | Rediscovered | Access | 
| Hybomorphus melanosomus | Saunders & Jekel, 1855 | Lord Howe Island ground weevil | June 1869 or before? | Lord Howe Island, New South Wales, Australia | Missing | Access | 
| Isotornus aterrimus | Wollaston, 1877 | ? | St. Helena | Missing | Access | |
| Leptopius etheridgei | (Olliff, 1889) | - | 1910's | Lord Howe Island, New South Wales, Australia | Missing | Access | 
| 
 Macrancylus linearis  | 
LeConte, 1876 | - | - | Carribean, Central America & North America | Erroneously listed as Extinct | Access | 
| Megacolabus sculpturatus | Broun, 1893 | Akaroa weevil | pre-1894 | Akaroa, Banks Peninsula, New Zealand | Missing | Access | 
| Neocnemis occidentalis | Crotch, 1867 | - | 1865 | Santa Maria, Azores | Extinct | Access | 
| Nesiobius barbatus | Wollaston, 1877 | - | 1880 | St. Helena | Extinct | Access | 
| Nesiobius breviusculus | Wollaston, 1877 | - | ? | St. Helena | Extinct | Access | 
| Nesiobius fimbriatus | Wollaston, 1887 | Fringed nesiobius weevil | ? | St. Helena | Extinct | Access | 
| Nesotocus kauaiensis | Perkins, 1900 | Kauai nesotocus weevil | ? | Kauai, Hawaiian Islands, USA | Missing | Access | 
| Ogasawarazo daitoensis | Voss, 1971 | - | (rediscovered) | Japan | Rediscovered | Access | 
| Oodemas laysanensis | Fullaway, 1914 | Laysan weevil | 1964 or June 1990 | Laysan, Midway, Nihoa and Necker Island, Hawaiian Islands, USA | Extinct | Access | 
| Orthorhinus lateralis | Pascoe, 1882 | - | ? | Lord Howe Island, New South Wales, Australia | Missing | Access | 
| Pentarthrum blackburni | Sharp, 1878 | Blackburn's weevil | 1961 | Hawaiian Islands, USA | Missing | Access | 
| Prothrombosternus tarsalis | Voss, 1965 | - | (rediscovered) | Rubeho Mountains, Tanzania | Rediscovered | Access | 
| 
 Pseudapocyrtus schadenbergi  | 
Heller, 1912 | - | (rediscovered) | northern Luzon, Philippines | Rediscovered | Access | 
| Rhyncogonus bryani | Perkins, 1919 | Bryan's weevil | 1911 | Laysan Hawaiian Islands, USA | Extinct | Access | 
| Rhyncogonus extraneus | Perkins, 1910 | - | 16 January 1941 | O'ahu, Hawaiian Islands, USA | Extinct | Access | 
| Rhyncogonus "tarakoi" | - | Tarakoi weevil | 1985 (1987?) | Tarakoi islet, Austral Islands | Missing | Access | 
| Thotmus halli | Broun, 1910 | Hall's weevil, Pitt Island weevil | 1906-1908 | Pitt Island, Chatham Islands, New Zealand | Extinct | Access | 
| Torishimazo minamiiwoensis | Morimoto, 2015 | - | (rediscovered) | Minami Iwo-jima, Japan | Rediscovered | Access | 
| Trigonoscuta rossi | Pierce, 1975 | Fort Ross weevil | 1975 | Fort Ross, California, USA | Missing | Access | 
| Trigonoscuta yorbalindae | Pierce, 1975 | Yorba Linda weevil | 1975 | Yorba Linda area, California, USA | Missing | Access | 
| Tychanopais sp. nov. 'Fred Cave' | - | Fred Cave tychanopais weevil | ? | Waitomo, North Island, New Zealand | Extinct | Access | 
| Tychiorhinus porrectus | Wollaston, 1877 | ? | St. Helena | Missing | Access | |
| Tymbopiptus valeas | Kuschel, 1987 | - | 1680 +/- 50 years BP | New Zealand | Extinct | Access | 
| Xestophasis nasalis | Wollaston, 1877 | ? | St. Helena | Missing | Access | |
| Xestophasis xerophilus | Decelle & Voss 1972 | - | pre-2003 | St. Helena | Missing | Access | 
| Xyleborus pele | Samuelson, 1981 | - | (rediscovered) | Hawaiian Islands, USA | Rediscovered | Access | 
| 
 Xyleborus pleiades 
 | 
Samuleson, 1981 | (rediscovered) | Maui & Moloka‘i, Hawaiian Islands, USA | Rediscovered | Access | 
Family: Dytiscidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Acilius sinensis | Peschet, 1915 | - | (rediscovered) | Yunnan Province, China | Extinct | Access | 
| Carabdytes plantaris | (Sharp, 1882) | - | (rediscovered) | New Zealand | Rediscovered | Access | 
| Coelhydrus brevicollis | Sharp, 1882 | - | (rediscovered) | South Africa | Rediscovered | Access | 
| Colymbetes minimus | Zaitzev, 1908 | (rediscovered) | China & Tibet | Rediscovered | Access | |
| Copelatus duodecimstriatus | Aubé, 1838 | - | (rediscovered) | Mascarene Islands | Rediscovered | Access | 
| Copelatus thiriouxi | Peschet, 1917 | - | (rediscovered) | Mascarene Islands | Rediscovered | Access | 
| Hydroporus compunctus | Wollaston, 1865 | - | ? | Canary Islands, North Africa | Missing | Access | 
| Hydroporus spangleri | Gordon, 1981 | Spangler's diving beetle, Spangler's hydroporus diving beetle | ? | Utah, USA | Missing | Access | 
| Hydroporus utahensis | Gordon, 1981 | Utah hydroporus diving beetle | ? | Utah, USA | Missing | Access | 
| Hydrotarsus pilosus | Guignot, 1949 | - | ? | Canary Islands, North Africa | Missing | Access | 
| Hygrotus artus | Fall, 1919 | Mono Lake diving beetle | (rediscovered) | Inyo County & Mono County, California, USA | Rediscovered | Access | 
| Hygrotus sylvanus | (Fall, 1917) | - | 1979 | USA | Extinct | Access | 
| Ilybius samokovi | Nilsson & Fery, 1993 | - | (rediscovered) | Rila Mountains, Bulgaria | Rediscovered | Access | 
| 
 Megadytes ducalis  | 
Sharp, 1882 | - | late 1800's | Bahia (southern), Brazil | Missing | Access | 
| Meridiorhantus orbignyi | (Balke, 1992) | - | (rediscovered) | Argentina & Brazil | Rediscovered | Access | 
| Paroster monteithi | (Brancucci, 1985) | (rediscovered) | New Caledonia | Rediscovered | Access | |
| Rhantus papuanus | Balfour-Browne, 1939 | - | 1919 | Papua New Guinea | Extinct | Access | 
| Siettitia balsetensis | Abeille de Perrin, 1904 | Perrin's cave beetle, Perrin's cave coleoptera | 1950's (1994?) | France | Extinct | Access | 
| Stictonectes canariensis | Machado, 1987 | - | ? | Canary Islands, North Africa | Missing | Access | 
| Stygoporus oregonensis | Larson and LaBonte, 1994 | (rediscovered) | Oregon, USA | Rediscovered | Access | 
Family: Elateridae (Click Beetles)
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Cardiophorus robustus | LeConte, 1853 | - | (rediscovered) | USA | Rediscovered | Access | 
| Dilobitarsus pendleburyi | Fleutiaux, 1934 | - | (rediscovered) | Malay Peninsula, Malaysia and Sumatra, Indonesia | Rediscovered | Access | 
Family: Elmidae (Rifle Beetles)
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Heterelmis stephani | Brown, 1972 | Stephan's riffle beetle, Stephan's heterelmis riffle beetle | 1993 | Arizona, USA | Extinct | Access | 
Family: Erirhinidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Echinocnemus reitteri | Schilsky, 1907 | - | 1904 | Israel | Missing | Access | 
| Echinocnemus sahlbergi | Schilsky, 1911 | - | 1904 | Israel | Missing | Access | 
Family: Eucnemidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Paleoxenus dohrni | Horn, 1878 | Dohrn's elegant beetle, Dohrn's elegant eucnemid beetle | ? | San Gabriel Mountains, California, USA | Extinct | Access | 
Family: Gyrinidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Metagyrinus sinensis | (Ochs, 1924) | (rediscovered) | China | Rediscovered | Access | 
Family: Haliplidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Algophilus lathridioides | Zimmermann, 1924 | Table Mountain hairy crawling water beetle | (rediscovered) | Simons Town, Cape Peninsula, South Africa | Rediscovered | Access | 
| Haliplus nitens | LeConte, 1850 | Disjunct crawling water beetle | 1932 | North America | Extinct | Access | 
Family: Hydraenidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Hydraena sappho | Janssens, 1965 | - | pre-1966 | Lefkada, Greece | Missing | Access | 
| Limnebius aridus | Perkins, 1980 | Animas (limnebius) minute moss beetle | 1952 | New Mexico, USA | Extinct | Access | 
| Limnebius asperatus | 
 Knisch, 1922  | 
- | (rediscovered) | northeastern Apennines, Italy | Rediscovered | Access | 
| Limnebius canariensis | d'Orchymont, 1938 | - | 6 June 1989 | Gran Canaria, Canary Islands, North Africa | Missing | Access | 
| Limnebius texanus | Perkins, 1980 | Texas (limnebius) minute moss beetle | ? | Texas, USA | Extinct | Access | 
| Limnebius utahensis | Perkins, 1980 | Utah (limnebius) minute moss beetle | 1947 | Utah, USA | Extinct | Access | 
| Ochthebius putnamensis | Blatchley, 1910 | Indiana (ochthebius) minute moss beetle | pre-1911 | Indiana, USA | Extinct | Access | 
| Sebasthetops omaliniformis | Jäch, 1998 | - | (rediscovered) | South Africa | Rediscovered | Access | 
Family: Hydrophilidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Allocotocerus mistus | d'Orchymont, 1939 | Table Mountain water scavenger beetle | ? | Table Mountain, Western Cape Province, South Africa | Missing | Access | 
| Protistolophus spangleri | Short, 2010 | - | (rediscovered) | Venezuela (southern) | Rediscovered | Access | 
Family: Hygrobiidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Hygrobia davidi | Bedel, 1883 | Chinese screech beetle, David's water hygrobia | 1873 (1883?) | Jiangxi Province, China | Extinct | Access | 
Family: Lampyridae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Photuris bethaniensis | McDermott, 1953 | Bethany Beach firefly | (rediscovered) | Sussex County, Delaware, USA | Rediscovered | Access | 
Family: Leiodidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Aglyptinus agathidioides | Blair, 1930 | Potters Bar beetle, Round fungus beetle | 14 April 1912 | Potters Bar, Hertfordshire, England | Extinct | Access | 
Family: Limnichidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Ersachus erichsonianus | Sharp, 1902 | - | (rediscovered) | Veracruz, Mexico | Rediscovered | Access | 
Family: Lucanidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Colophon thunbergi | Westwood, 1855 | - | (rediscovered) | Cape Province, South Africa | Rediscovered | Access | 
| 
 Lissotes crenatus 
 | 
(Westwood, 1855) | - | (rediscovered) | Tasmania, Australia | Rediscovered | Access | 
| 
 Sclerostomulus nitidus 
 | 
(Benesh, 1955) | (rediscovered) | Cerro Poqui, Chile | Rediscovered | Access | 
Family: Meloidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Tetraolytta gerardi | (Pic, 1919) | (rediscovered) | Brazil | Rediscovered | Access | 
Family: Melolonthidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Apicencya calva | (Waterhouse, 1882) | 108+ years ago | Madagascar | Missing | Access | |
| Bothynus cribrarius | (Fairmaire, 1878) | - | (rediscovered) | Atlantic Forest, Brazil | Rediscovered | Access | 
Family: Melyridae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Trichochrous kernensis | (Blaisdell, 1926) | - | (rediscovered) | California, USA | Rediscovered | Access | 
Family: Mordellidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Tomoxia howensis | Lea, 1917 | - | 1880's | Lord Howe Island, New South Wales, Australia | Missing | Access | 
Family: Nitidulidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| 
 Apetasimus atratus 
 | 
(Scott in Sharp and Scott, 1908) | - | 1894 or later | Maui, Hawaiian Islands, USA | Missing | Access | 
| Apetasimus expers | (Blackburn in Blackburn and Sharp, 1885) | - | ? | Maui, Hawaiian Islands, USA | Missing | Access | 
| Apetasimus guttatus | (Sharp, 1881) | - | ? | Hawaiian Islands, USA | Missing | Access | 
| Apetasimus kauaiensis | (Scott in Sharp & Scott, 1908) | - | 1894 or later | Kauai, Hawaiian Islands, USA | Missing | Access | 
| Apetasimus sordidus | (Sharp, 1881) | - | ? | Hawaiian Islands, USA | Missing | Access | 
| Apetasimus subalatus | (Scott in Sharp & Scott, 1908) | - | ? | Hawaiian Islands, USA | Missing | Access | 
| Meligethes salvan | Audisio, De Biase & Antonini, 2003 | - | (rediscovered) | Italy | Rediscovered | Access | 
| Nesopeplus serratus | Scott, 1908 | Souring beetle | c. 40 years ago | Hawaiian Islands, USA | Missing | Access | 
Family: Psephenidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Mataeopsephus nitidipennis | Waterhouse, 1876 | (rediscovered) | China | Rediscovered | Access | 
Family: Rutelidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Balanogonia constricta | Paucar-Cabrera | (rediscovered) | Mexico | Rediscovered | Access | 
Family: Scarabaeidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Ahermodontus bischoffi | (Všetečka, 1939) | - | (rediscovered) | Albania | Extinct | Access | 
| 
 Anomala schaefferi  | 
Seidel, 2021 | Padre Island tibial scarab, Tibial scarab | 19 May 1976 | Padre Island, Texas, USA | Missing | Access | 
| Anomala tibialis | Schaeffer, 1906 | Padre Island tibial scarab, Tibial scarab | 19 May 1976 | Padre Island, Texas, USA | Invalid (junior homonym) | Access | 
| Anomis vulpicolor | (Meyrick, 1928) | Red Anomis noctuid moth, Fox-coloured owlet moth | ? | Hawai’i, Moloka’i, and O’ahu, Hawaiian Islands, USA | Missing | Access | 
| 
 Apogonia insulana  | 
Karsch, 1882 | (rediscovered) | Príncipe | Rediscovered | Access | |
| Coenonycha globosa | McClay, 1943 | - | (rediscovered) | Washington, USA | Rediscovered | Access | 
| Copris pristinus | Pierce, 1946 | - | Late Pleistocene | Los Angeles, California, USA | Extinct | Access | 
| Eodrepanus coopei | Barbero, Palestrini & Roggero, 2009 | - | c.118,000 yBP | England | Extinct | Access | 
| Eophileurus convexus | (Arrow in Andrews, 1900) | - | ? | Christmas Island, Australia | Missing | Access | 
| Hoplia walterrossii | Sabatinelli in Baraud, 1992 | (rediscovered) | Molise, Italy (southern) | Rediscovered | Access | |
| Onthophagus everestae | Pierce, 1946 | - | Late Pleistocene | Los Angeles, California, USA | Extinct | Access | 
| Onthophagus pilauco | Tello et al., 2021 | ~15.2 k cal yr BP | Chile | Extinct | Access | |
| Oryctes chevrolatii | Guérin-Méneville, 1844 | ? | Mauritius, Mascarene Islands | Extinct | Access | |
| Oryctes minor | Waterhouse, 1876 | ? | Rodrigues, Mascarene Islands | Extinct | Access | |
| Oryctes tarandus | (Olivier, 1789) | ? | Mauritius, Mascarene Islands | Extinct | Access | |
| Paracryptocanthon borgmeieri | Vulcano, Pereira & Martinez, 1976 | - | (rediscovered) | Brazil | Rediscovered | Access | 
| Phanaeus violetae | Zunino, 2013 | Late Pleistocene | Ecuador | Extinct | Access | |
| Prodontria sp. nov. 'New Zealand' | - | - | ? | New Zealand | Extinct | Access | 
| Scarabaeus cancer | (Arrow, 1919) | - | (rediscovered) | Kuando Kubango Province, Angola | Rediscovered | Access | 
| 
 Scarabaeus sevoistra  | 
Alluaud, 1902 | - | (rediscovered) | southern Madagascar | Rediscovered | Access | 
| 
 Scybalophagus brellenthinae  | 
Tello et al., 2023 | - | ~16 400 and ~12 800 cal a bp. | Chile | Extinct | Access | 
| 
 Semanopterus kingstoni  | 
Reid & Tees, 2023 | - | 30 January 1979 | Mt. Gower, Lord Howe Island, New South Wales, Australia | Missing | Access | 
| Stenosternus costatus | Karsch, 1881 | - | (rediscovered) | São Tomé, Gulf of Guinea, West Africa | Rediscovered | Access | 
Family: Silphidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Nicrophorus hispaniola | Sikes & Peck, 2000 | - | (rediscovered) | Dominican Republic, Hispaniola | Rediscovered | Access | 
Family: Staphylinidae (Rove Beetles)
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Aleochara repetita | Sharp, 1887 | (rediscovered) | Brazil & Panamá | Rediscovered | Access | |
| Batriasymmodes parki | Barr, 1987 | Antloving beetle | ? | Neely Farm Cave, Mercer County, West Virginia, USA | Extinct | Access | 
| Batrisodes profundus | Park, 1956 | Profound Cave rove beetle | ? | Alabama, USA | Missing | Access | 
| Darwinilus sedarisi | Chatzimanolis, 2018 | - | Between 6 September 1832 and 17 October 1832 | Bahía Blanca, Argentina | Missing | Access | 
| Geostiba melanocephala | (Crotch, 1867) | 1867 or before | Azores, Macaronesia | Missing | Access | |
| Hesperus gigas | (Lea, 1929) | - | 1910's | Mount Lidgbird, Lord Howe Island, New South Wales, Australia | Extinct | Access | 
| 
 Losiusa angusticollis  | 
Seevers, 1978 | - | (rediscovered) | Massachusetts, USA | Rediscovered | Access | 
| Mastopsenius australis | Seevers, 1945 | - | (rediscovered) | Northern Territory & Queensland, Australia | Rediscovered | Access | 
| Menoedius andrewesi | 
 Fauvel, 1903  | 
- | (rediscovered) | Nilgiri Mountains, Tamil Nadu, India | Rediscovered | Access | 
| Oxyporus flohri | Sharp, 1887 | - | (rediscovered) | Guatemala & Veracruz, México | Rediscovered | Access | 
| Paraneseuthia holzneri | (Franz, 1976) | - | (rediscovered) | Japan | Rediscovered | Access | 
| 
 Phytosus schatzmayri  | 
Bernhauer, 1941 | 1935 | São Miguel Island, Azores, Macaronesia | Missing | Access | |
| 
 Scydmaenus kasuganus  | 
Franz, 1976 | (rediscovered) | Japan (including Mount Kasuga ad Nara) | Rediscovered | Access | 
Family: Tenebrionidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Alphasida nitida | Bedel, 1918 | - | (rediscovered) | Morocco | Rediscovered | Access | 
| Asida septemsis | (Pérez Vera, Ruiz & Avila, 2012) | - | (rediscovered) | Morocco | Rediscovered | Access | 
| Coniontis remnans | Pierce, 1954 | - | Late Pleistocene | Rancho La Brea Tar Pit, California, USA | Missing | Access | 
| Helenomelas basilewskyi | Ardoin, 1972 | - | 1960's | St. Helena | Extinct | Access | 
| Hymenophorus candeli | (Pardo-Alcaide, 1978) | - | (rediscovered) | Morocco | Rediscovered | Access | 
| Nesotes azoricus | (Crotch, 1867) | - | 1930 | São Miguel Island, Azores | Extinct | Access | 
| Penthicoides seriatoporus | Fairmaire, 1896 | - | (rediscovered) | Karnataka, India | Rediscovered | Access | 
| Tarphiophasis wollastoni | Ardoin, 1972 | - | pre-2003 | St. Helena | Missing | Access | 
| Terametus capicola | Motschulsky, 1869 | (rediscovered) | South Africa | Rediscovered | Access | 
Family: Trogidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Trox youngai | Strümpher & Scholtz, 2011 | - | ? | South Africa | Missing | Access | 
Family: Trogossitidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| 
 Tenebroides atiu 
 | 
Kolibáč & Porch, 2020 | - | c. 1400 yBP | Atiu, Cook Islands | Extinct | Access | 
| Tenebroides mihiura | Kolibáč & Porch, 2020 | - | c. 2000-2500 yBP | Tubuai, Austral Islands, French Polynesia | Extinct | Access | 
| Tenebroides moorea | Kolibáč & Porch, 2020 | - | c. 3000-4000 yBP | Mo’orea, Society Islands, French Polynesia | Extinct | Access | 
| Tenebroides raivavae | Kolibáč & Porch, 2020 | - | c. 1000-1500 yBP | Raivavae, Austral Islands, French Polynesia | Extinct | Access | 
| Tenebroides rimatara | Kolibáč & Porch, 2020 | - | c. 3500 yBP | Rimatara, Austral Islands, French Polynesia | Extinct | Access | 
| Tenebroides tubuai | Kolibáč & Porch, 2020 | - | c. 2000-2500 yBP | Tubuai, Austral Islands, French Polynesia | Extinct | Access | 
Family: Ulodidae
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Waitomophylax worthyi | (Leschen & Rhode, 2002) | - | Holocene | New Zealand | Extinct | Access | 
Family: Zopheridae (Ironclad Beetles)
| Scientific Name | Author/s | Common Name | Last Record | Distribution | Status | Taxon Profile | 
| Antilissus makauwahi | Porch, 2020 | subfossil | Kauai, Hawaiian Islands, USA | Extinct? | Access | |
| Pycnomerus mahanatoa | Porch et al., 2020 | subfossil | Raivavae, French Polynesia | Extinct? | Access | |
| Pycnomerus prebblei | Porch & Smith, 2017 | - | subfossil | Rimatara, French Polynesia | Extinct | Access | 
| Pycnomerus rairua | Porch et al., 2020 | subfossil | Raivavae, French Polynesia | Extinct? | Access | |
| Pycnomerus raivavae | Porch et al., 2020 | subfossil | Raivavae, French Polynesia | Extinct? | Access | |
| Pycnomerus rimatara | Porch & Smith, 2017 | - | subfossil | Rimatara, French Polynesia | Extinct | Access | 
| Pycnomerus taralewisae | Porch et al., 2020 | subfossil | Raivavae, French Polynesia | Extinct? | Access | |
| 
 Pycnomerus vavitu 
 | 
Porch et al., 2020 | subfossil | Raivavae, French Polynesia | Extinct? | Access | 
<< Back to the Recently Extinct Animals database
Appendix 1: Beetles with unknown type locality
Barr, Cheryl B. and Shepard, William D. (2020). Hiding in plain sight: rediscovery and review of Parygrus Erichson, 1847, with description of five new species from the Neotropics (Coleoptera: Byrrhoidea: Dryopidae). Zootaxa 4755(1): 99-128. https://doi.org/10.11646/zootaxa.4755.1.4 [Parygrus erichsoni from Colombia]
Appendix 2: Species erroneously listed as extinct
"Incidentally, one of these six species is Carabdytes novaecaledoniae (Balfour-Browne) (included in the Red List since 1996 as Rhantus novacaledoniae), which is not Extinct and has been recorded recently from New Caledonia (Jäch & Balke, 2008)"
Source: Cowie, Robert H., Bouchet, Philippe and Fontaine, Benoît. (2022). The Sixth Mass Extinction: fact, fiction or speculation? Biological Reviews 97: 640-663. doi: 10.1111/brv.12816