Heterelmis stephani Brown, 1972

Stephan's riffle beetle, Stephan's heterelmis riffle beetle



Taxonomy & Nomenclature



Conservation Status

Last record: 1993 (Barr et al. 1993?)



Arizona, USA


Biology & Ecology










Original scientific description:

Brown, Harley P. (1972). Synopsis of the Genus Heterelmis Sharp in the United States, with description of a new species from Arizona (Coleoptera, Dryopoidea, Elmidae). Entomological News 83: 229-238.


Other references:

Barr, C. B. (1991). Personal letter to Robert A. Johnson regarding the distribution and status of Heterelmis stephani, dated December, 19(1991), 2.

Barr, C. B. and Shepard, W. D. (1993). Survey for Heterelmis stephani Brown (Insecta: Coleoptera: Elmidae) in Madera Canyon and other localities in the Santa Rita Mountains, Arizona. Final Report prepared for US Forest Service.

Bosse, L. S., Tuff, D. W. and Brown, H. P. (1988). A new species of Heterelmis from Texas (Coleoptera: Elmidae). Southwestern Naturalist 33: 199-203.

Brown, Harley P. and Shepard, William D. (1979). Type Specimens in the Zoological Collections of the Stovall Museum of Science and History. The Southwestern Naturalist 24(1): 195-197.

Carlton, Christopher E. et al. (2005). Obituary and dedication: Karl Heinz Stephan 1931-2005. The Coleopterists Bulletin, 59(3): 277-283.

Gonzales, Tina K. (2008). Conservation genetics of the Comal Springs riffle beetle (Heterelmis comalensis) populations in central Texas, with examination of molecular and morphological variation in Heterelmis sp. throughout Texas. Master of Science Thesis, Texas State University, San Marcos.

Whiteman, Noah K. and Sites, Robert W. (2008). Aquatic insects as umbrella species for ecosystem protection in Death Valley National Park. Journal of Insect Conservation 12: 499-509.


<< Back to the Coleoptera (Beetles) database