Copris pristinus Pierce, 1946:124



Taxonomy & Nomenclature



Conservation Status


Last record: Late Pleistocene



Los Angeles, California, USA












Original scientific description:

Pierce, W. D. (1946). Descriptions of the dung beetles (Scarabaeidae) of the tar pits. Bulletin, Southern California Academy of Sciences 45: 119-131.


Other references:

Halffter, G. (1959). Etologia y paleontologia de Scarabaeinae (Coleoptera, Scarabaeidae). Ciencia 19, 165-178. [p. 176]

Krell, Frank-Thorsten. (2000). The fossil record of Mesozoic and Tertiary Scarabaeoidea (Coleoptera: Polyphaga).
Invertebrate Taxonomy 14: 871-905. [p. 879]

Krell, Frank-Thorsten. (2004). Catalogue of fossil Scarabaeoidea (Coleoptera: Polyphaga) of the Mesozoic and Tertiary. Unpublished. 61 pp. [accessed 30 March 2014]

Krell, Frank-Thorsten. (2006). Fossil Record and Evolution of Scarabaeoidea (Coleoptera: Polyphaga). Coleopterists Society Monograph 5 (Supplement to The Coleopterists Bulletin 60): 120-143. [p. 132]

Krell, Frank-Thorsten. (2007). Catalogue of fossil Scarabaeoidea (Coleoptera: Polyphaga) of the Mesozoic and Tertiary - Version 2007 -. Denver Museum of Nature & Science Technical Report 2007-8: 79 pp.

Matthews, E. G. (1961). A revision of the genus Copris Müller of the Western Hemisphere (Coleoptera, Scarabaeidae). Entomologica Americana, New Series 41: 1-139. [p. 35,67,69]

Matthews, E. G., and Halffter, G. (1968). New data on American Copris with discussion of a fossil species. Ciencia 26: 147-162. [p. 160]

Miller, S. E. (1997). Late Quaternary insects of Rancho La Brea, California, U.S.A: in A. C. Ashworth, P. C. Buckland, and J. P. Sadler, eds., Studies in Quaternary Entomology - an inordinate fondness for insects: Quaternary Proceedings 5: 185-191. [p. 188]

Miller, S. E., Gordon, R. D. and Howden, H. F. (1981). Reevaluation of Pleistocene scarab beetles from Rancho La Brea, California (Coleoptera, Scarabaeidae): Proceedings of the Entomological Society of Washington 83: 625-630. [p. 626]

Morón, M.Á. (2003). Antecedentes. In ‘Atlas de los escarabajos de México. Coleoptera: Lamellicornia Vol. II. Familias Scarabaeidae, Trogidae, Passalidae y Lucanidae’ (ed. M.Á. Morón) pp. 11-18 (Argania edition: Barcelona). [p. 12]

Sphon, G. G. (1973). Additional type specimens of fossil Invertebrata in the collections of the Natural History
Museum of Los Angeles County. Contributions in Science, Natural History Museum, Los Angeles County
250, 75 pp. [p. 52]

Stock, C., and Harris, J.M. (1992). ‘Rancho La Brea. A Record of Pleistocene life in California.’ 7th Ed. (Natural
History Museum of Los Angeles County: Los Angeles.) xiv, 113 pp. [p. 70,84]


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