Horologion speokoites Valentine, 1932:3

Arbuckle Cave ground beetle



Taxonomy & Nomenclature

Synonym/s: Horologion speokites Valentine, 1932 [orth. error]


Conservation Status


Last record: 12 July 1931 (Harden et al., 2024)



Arbuckle Cave, West Virginia (southeastern), USA


Biology & Ecology




Only known from the holotype: USNM 44255 (male)






Original scientific description:

Valentine, J. M. (1932). Horologion, a new genus of cave beetles (fam. Carabidae). Annals of the Entomological Society of America 25: 1-11


Other references:

Barr Jr TC (1994) Obituary. J. Manson Valentine. NSS News 52: 355.

Bosquet, Yves. (2012). Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico. ZooKeys 245: 1-1722. [p. 711]

Davies WE (1965) Caverns of West Virginia. West Virginia Geological and Economic Survey Vol. 19A, [vi +] 330 pp.

Erwin, Terry, Donald R. Whitehead, and George E. Ball / Arnett, Ross H., ed. (1977). Checklist of the Beetles of North and Central America and the West Indies. Family 4. Carabidae, The Ground Beetles. Kinderhook, New York, USA: The Biological Research Institute of America, Inc.

Fong DW, Culver DC, Hobbs III HH, Pipan T (2007) The Invertebrate Cave Fauna of West Virginia, 2nd Edn. West Virginia Speleological Survey, Bulletin # 16, 163 pp.

Harden, Curt W., Davidson, Robert L., Malabad, Thomas E., Caterino, Michael S. and Maddison, David R. (2024). Phylogenetic systematics of the enigmatic genus Horologion Valentine, 1932 (Coleoptera, Carabidae, Trechinae, Horologionini), with description of a new species from Bath County, Virginia. Subterranean Biology 48: 1-49.


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