Paralouatta varonai Rivero & Arredondo, 1991:1

Galeran monkey, Cuban monkey



Taxonomy & Nomenclature



Conservation Status


Last record: Quaternary









Holotype: MNHNH V 194 (ex. MNHNH 25-90)

Other material:

MNHNCu 76.1010–1013
MNHNCu 76.1024
MNHNCu 76.1035






Original scientific description:

Rivero, M. and Arredondo, Oscar. (1991). Paralouatta varonai, a new Quaternary platyrrhine from Cuba. Journal of Human Evolution 21: 1-11. [Abstract]


Other references:

Alcover, Josep Antoni et al. (1998). Mammal Species of the World: Additional Data on Insular Mammals. American Museum Novitates 3248, 29 pp., 1 table.

Beck, Robin M. D. et al. (2023). Total evidence phylogeny of platyrrhine primates and a comparison of undated and tip-dating approaches. Journal of Human Evolution 174: 103293.

Borroto-Páez, Rafael and Mancina, Carlos A. (2017). Biodiversity and conservation of Cuban mammals: past, present, and invasive species. Journal of Mammalogy 98(4): 964-985.

Borroto-Páez, Rafael, Mancina, Carlos A., Woods, Charles A. and Kilpatrick, C. William. (2012). Checklist: Updated Checklist of Endemic Terrestrial Mammals of the West Indies, pp. 389-415. In: Borroto-Páez, Rafael, Woods, Charles A. and Sergile, F. E. (eds.). Terrestrial Mammals of the West Indies: Contributions. Gainesville, Florida: Florida Museum of Natural History and Wacahoota Press. 482 pp.

Herrera-Uria, Jane et al. (2015). Type specimens housed in the National Museum of Natural History of Cuba. Solenodon 12: 84-123.

Horovitz, I. and MacPhee, R. D. E. (1999). The Quaternary Cuban platyrrhine Paraloutta varonai and the origin of Antillean monkeys. Journal of Human Evolution 36: 33-68.

Ignacio Ramos García and Rafael Borroto Páez. (2000). Ejemplares tipo de mamíferos en colecciones cubanas. Orsis 15: 75-89.

MacPhee, R. D. E. (1993). From Cuba: A mandible of Paralouatta. Evolutionary Anthropology 2(2): 42.

MacPhee, R.D.E. & Horovitz, I. 2004 New craniodental remains of the Quaternary Jamaican monkey Xenothrix mcgregori (Xenotrichini, Callicebinae, Pitheciidae), with a reconsideration of the Aotus hypothesis. American Museum Novitates 3434, 1-51.

MacPhee, R. D. E. and Meldrum, Jeff. (2006). Postcranial Remains of the Extinct Monkeys of the Greater Antilles, with Evidence for Semiterrestriality in Paralouatta. American Museum Novitates, No. 3516: 1-65.

Sivault, Elise et al. (2023). Can body mass and skull morphology predict seed and fruit ingestion potential for mammal species? A test using extant species and its application to extinct species. Functional Ecology.

Turvey, Samuel T. (2009). Holocene mammal extinctions, pp. 41-61. In: Turvey, Samuel T. (ed.). Holocene Extinctions. Oxford, UK & New York, USA: Oxford University Press. xii + 352 pp.

Turvey, Samuel T. and Fritz, Susanne A. (2011). The ghosts of mammals past: biological and geographical patterns of global mammalian extinction across the Holocene. Philosophical Transactions of the Royal Society B 366(1577): 2564-2576. [Supplementary Information]

Upham, Nathan S. (2017). Past and present of insular Caribbean mammals: understanding Holocene extinctions to inform modern biodiversity conservation. Journal of Mammalogy 98(4): 913-917.


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