Pachylemur jullyi (G. Grandidier, 1899)

Northern giant ruffed lemur



Taxonomy & Nomenclature

Synonym/s: Palaeochirogalus jullyi G. Grandidier, 1899; Lemur jullyi Standing, 1904; Lemur maxiensis Standing, 1904; Lemur majori Standing, 1908


Conservation Status
















Original scientific description:

Grandidier, G. (1899). Description d'ossements de Lémuriens disparus. Bulletin du Muséum national d'histoire naturelle. 5: 344–348.


Other references:

Alcover, Josep Antoni et al. (1998). Mammal Species of the World: Additional Data on Insular Mammals. American Museum Novitates 3248, 29 pp., 1 table.

Crovella, S., Montagnon, D., Rakotosamimanana, B. & Rumpler, Y. 1994. Molecular biology and systematics of an extinct lemur: Pachylemur insignis. Primates 35, 519-522.

Crowley, Brooke Erin, Godfrey, Laurie Rohde and Samonds, Karen Elizabeth. (2023). What can hippopotamus isotopes tell us about past distributions of C4 grassy biomes on Madagascar? Plants, People, Planet.

Joseph, Grant S. and Seymour, Colleen L. (2022). Are Madagascar's obligate grazing-lawns ancient and evolved with endemic herbivores, or recently selected by introduced cattle? Biology Letters 18: 20220212.

Jungers, W.L., Demes, B. & Godfrey, L.R. 2008 How big were the "giant" extinct lemurs of Madagascar? In Elwyn Simons: a search for origins (ed. J.G. Fleagle & C.C. Gilbert), pp. 343-360. New York: Springer.

Karanth, K.P., Delefosse, T., Rakotosamimanana, B., Parsons, T.J. & Yoder, A.D. 2005. Ancient DNA from giant extinct lemurs confirms single origin of Malagasy primates. Proceedings of the National Academy of Sciences of the USA 102, 5090-5095.

Samonds, Karen E. et al. (2019). A new late Pleistocene subfossil site (Tsaramody, Sambaina basin, central Madagascar) with implications for the chronology of habitat and megafaunal community change on Madagascar’s Central Highlands. Journal of Quaternary Science 34(6): 379-392.

Standing, H.F. (1904). "Rapport sur des ossements sub-fossiles provenant d'Ampasambazibma. 1. — Les Lémuriens (Suite)". Bulletin de l'Académie malgache. 3: 305–310.

Standing, H.F. (1908). On recently discovered subfossil primates from Madagascar. Transactions of the Zoological Society of London. 18 (2): 59–162.

Turvey, Samuel T. (2009). Holocene mammal extinctions, pp. 41-61. In: Turvey, Samuel T. (ed.). Holocene Extinctions. Oxford, UK & New York, USA: Oxford University Press. xii + 352 pp.

Turvey, Samuel T. and Fritz, Susanne A. (2011). The ghosts of mammals past: biological and geographical patterns of global mammalian extinction across the Holocene. Philosophical Transactions of the Royal Society B 366(1577): 2564-2576. [Supplementary Information]


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