Stephanorhinus kirchbergensis Jäger, 1839
Merck's rhinoceros
Taxonomy & Nomenclature
Synonym/s: Rhinoceros merckii Jäger, 1839; Rhinoceros mercki Jäger, 1839; Dicerorhinus merckii Jäger, 1839; Dicerorhinus orientalis; Stephanorhinus orientalis; See (Billia, 2008; Billia & Petronio, 2009) for a complete synonymy
Conservation Status
Extinct
Last record: Late Pleistocene
Distribution
Eurasia
Biology
Hypodigm
SMNK PAL 4254 (Pandolfi, 2021)
SMNS 6617.2.12.67.3 (Pandolfi, 2021)
Media
References
Original scientific description:
Jäger, G. F. (1835-39). Über die fossilen Säugetiere welche in Würtemberg in verschiedenen Formationen aufgefunden worden sind, nebst geognotischen Bemerkungen über diese Formationen. C. Erhard Verlag, 139 pp., Stuttgart.
Other references:
Abbott W.J.L. (1890). Notes on some Pleistocene sections in and near London. Proceedings of the Geologists’ Association, 11: 473-480.
Adam K-D. (1958). Dicerorhinus kirchbergensis (Jäger) aus einer Karsthöhle bei Crni Kal (Istrien, Jugoslawien) [with Slovenian abstract]. Razprave [Dissertationes], Academia Scientiarum et Artium Slovenica (Slovenska Akademija Znanosti i Umetnosti [SAZU]), 4 (4): 437-440.
Alekseeva E.V. (1980). Mlekopitayushchye Pleystozena YugoVostoka Zapadnoy Sibiri (Hishchnye, Hobotnye, Kopytnye) [Pleistocene Mammals from southeast of Western Siberia]. AN SSSR, Dal’nevostochny Nauchny Zentr Biologii, Izd-vo “Nauka”, 187 pp., Moskva [in Russian].
Irena Axmanová, Jan Robovský, Lubomír Tichý, Jiří Danihelka, Elena Troeva, Albert Protopopov and Milan Chytrý. (2020). Habitats of Pleistocene megaherbivores reconstructed from the frozen fauna remains. Ecography 43: 1-11.
Bar-Yossef O. & Chernov E. (1972). On the Palaeo-Ecological History of the Site of ’Ubeidiya. Israel Academy of Human Sciences and Humanities, 35 pp., Jerusalem.
Belyaeva E.I. (1935). Nekotorye dannye o chetvertichnykh mlekopitayuscikh iz Nizhnevolzhskogo kraya po materialam muzeya g. Pugach. Zemlya [Some data on the Quaternary mammals from the Lower Volga area based on the material in the Pugachev Region Municipal Museum]. Trudy Komissii po Izucheniyu Chetvertichnogo Perioda, 4: 303-308, Moskva [in Russian, German summ.].
Belyaeva E.I. (1939). Ob ostatkakh iskopaemogo nosoroga iz okrestnostey g. Rybinska. [On fossil rhinoceros remains from the environs of Rybinsk]. In Belyaeva E.I. (ed.), Voprosy Stratigrafii Chetvertichnykh Otlozhenii i ee Paleontologicheskoe Obosnovanie. Byulleten’ Komissii po Izucheniyu Chetvertichnogo Perioda, 5: 69-90, Izd-vo AN SSSR, Moskva/Leningrad [in Russian].
Belyaeva E.I. (1940). Novye nakhodki ostatkov nosoroga Merka na territorii SSSR [New finds of Merck’s rhinoceros remains on Soviet Union territory]. Priroda (Paleozoologya), 8: 82, Izd-vo AN SSSR [in Russian].
Bernsen, J. J. A. (1927). The geology of the Tegelian Clay and its fossil remains of Rhinoceros. Thesis, University of Amsterdam, Teulings’ Koninkl. Drukkerrijen, 108 pp., ‘s-Hertogenbosch.
Billia, Emmanuel M. E. (2007). Revision of the fossil material attributed to Stephanorhinus kirchbergensis (Jäger, 1839) (Mammalia, Rhinocerotidae) preserved in the museum collections of the Russian Federation. In Mironov A.G., Catto N.R. & Erbajeva M.A. (eds.), Proceedings of the INQUA International Symposium “Quaternary of Baikalia - Stratigraphy, Paleontology and Paleoenvironments of the Pliocene-Pleistocene of Transbaikalia and Interregional Correlations”. Quaternary International 179(1): 25-37.
Billia, E.M.E., 2008. Revision of the fossil material attributed to Stephanorhinus kirchbergensis (Jäger 1839) (Mammalia, Rhinocerotidae) preserved in the museum collections of the Russian Federation. Quat. Int. 179, 25–37.
Billia E.M.E. 2011 Occurrences of Stephanorhinus kirchbergensis (Jager, 1839) (Mammalian, Rhinocerotidae) in Eurasia - An account. Acta Palaeontologica Romaniae 7, 17-40.
Billia, Emmanuel M. E. and Petronio, Carmelo. (2009). Selected records of Stephanorhinus kirchbergensis (Jäger, 1839) (Mammalia, Rhinocerotidae) in Italy. Bollettino della Società Paleontologica Italiana 48(1): 21-32.
Borsuk-Bialynicka M. & Jakubowski G. (1972). The skull of Dicerorhinus mercki (Jäger) from Warsaw. Prace Muzeum Ziemi [Proceedings of the Museum of the Earth], 20: 187-199, Polska Akademia Nauk (Prace Paleozoologiczne), Warszawa.
Brandt, J. F. (1877). Versuch einer Monographie der Tichorhinen Nashörner nebst Bemerkungen über Rhinoceros leptorhinus Cuvier u.s.w. Mémoires de l’Academie Impériale des Sciences de St.-Pétersbourg, s. 7, 24 (4): 1-135.
Briatico, Giuseppe and Bocherens, Hervé. (2023). Middle Pleistocene ecology in central Italy. New isotopic insights from fauna tooth enamel of Casal de’ Pazzi (Rome, Italy). Journal of Mediterranean Earth Sciences. doi:10.13133/2280-6148/18031
Burkanova, Elena M. et al. (In Press, 2020). Stephanorhinus kirchbergensis (Jäger, 1839) (Mammalia, Rhinocerotidae) from the Po valley (Lombardia, Northern Italy): Possible diet/nutrition and living conditions. Quaternary International. https://doi.org/10.1016/j.quaint.2020.07.031 [Abstract]
Cappellini, Enrico et al. (2019). Early Pleistocene enamel proteome from Dmanisi resolves Stephanorhinus phylogeny. Nature. DOI: https://doi.org/10.1038/s41586-019-1555-y [Abstract]
Chen, Shaokun, Pang, Libo, Yan, Yaling, Wei, Guangbiao and Yue, Zongying. (Accepted, 2021). First discovery of Dicerorhinus sumatrensis from Yanjinggou provides insights into the Pleistocene Rhinocerotidae of South China. Acta Geologica Sinica. https://doi.org/10.1111/1755-6724.14719 [Abstract]
Chen, Xi, Wu SM, Wang P, et al. (2020). A report on the Late Pleistocene vertebrate fossils from the Zhangshan locality, Suqian, Jiangsu Province. Acta Anthropologica Sinica 39(2): 319-331.
Chersky I.D. (1874). Opisanie cherepa nosoroga, razlichnago ot’ Rhinoceros tichorhinus [Description of a rhinoceros skull different from Rhinoceros tichorhinus]. Zapiski Imperatorskoy Akademii Nauk”, 25: 65-75 [in Russian].
Choe, R. S., Han, K. S., Kim, S. C., U, C., Ho, C. U., Kang, I. (2020). Late Pleistocene fauna from Chongphadae Cave, Hwangju County, Democratic People’s Republic of Korea. Quaternary Research [2020]: 1-13. https://doi.org/10.1017/qua.2020.9 [Abstract]
Czyzewska T. (1962). Uzebienie górnej szczeki Dicerorhinus mercki (Jäger) ze Szczesliwice kolo Warszawy [Upper dentition of Dicerorhinus mercki (Jäger) from Szczesliwice near Warszawa (Poland)]. Acta Paleontologica Polonica, 7 (1-2): 223-234 [in Polish, Russian and English abstracts].
Diedrich, Cajus G. (2023). Extinct Eurasian rhinoceros Coelodonta and Stephanorhinus dental pathologies and tooth change modus. Quaternary Science Reviews 301: 107922. https://doi.org/10.1016/j.quascirev.2022.107922
Dong, Wei, Xu, Qinqi, Jin, Changzhu and Liu, Jinyi. (1999). Quaternary herbivore fauna in northeastern China: Evolution under climatic change. Chinese Science Bulletin 44(Suppl.): 129-132.
Dong, Wei and Zhang, Yingqi. (2022). Biochronologic sequences of the Pleistocene mammalian faunas in China and correlations with numeric dating. Quaternary Sciences 42(5): 1227-1246. https://doi.org/10.11928/j.issn.1001-7410.2022.05.01
Döppes D. & Rabeder G. (1997). Pliozäne und Pleistozäne Faunen Österreichs - Ein Katalog der wichtigsten Fossilfundstellen und ihrer Faunen. Mitteilungen der Kommission für Quartärforschung der Österreichischen Akademie der Wissenschaften: 69.
Dubrovo I.A. (1957). Ob ostatkakh Parelephas wüsti (M. Pavl.) i Rhinoceros mercki Jäger iz Yakutii [On remains of Parelephas wüsti (M. Pavl.) and Rhinoceros mercki Jäger from Yakutya]. Byulleten’ Komissii po Izucheniyu Chetvertichnogo Perioda, 21: 97-104, Izd-vo AN SSSR, Moskva/Leningrad [in Russian].
Flerov K.K., Trominov B.A. & Yanoskaya N.M. (1955). Istorya Fauny Mlekopitayushchikh v Chetvertichnom Periode [History of Mammalian Fauna in the Quaternary Period]. AN SSSR (Geologichesky Muzey): 5-39, Moskva [in Russian].
Fortelius, Mikael, Mazza, Paul and Sala, Benedetto. (1993). Stephanorhinus (Mammalia: Rhinocerotidae) of the western European Pleistocene, with a revision of S. etruscus (Falconer, 1868). Plaeontographia Italica 80: 63-155.
Freudenberg W. (1914). Die Säugetiere des Altern Quartärs von Mitteleuropa mit besonderes Berücksichtigung der Faunen von Hundsheim und Deutsch-Altenburg in Niederösterreich nebst Bemerkungen über verwandte Formen anderer Fundorte. Geologische und Paläontologische Abhandlungen, NF 12 (4-5): 455-671.
Fu Zheni (2002). Chetvertichnaya fauna mlekopitayushchikh Pleystozena Dunbeya (KNR) i ego spezifika [Quaternary mammalian fauna of the Pleistocene of Dumbey (P.R. of Korea) and its detailed note]. Arkheologya, Etnografya i Antropologya Evrazii 1(9): 6-15 [in Russian].
Gliozzi E., Abbazzi L., Ambrosetti P.G., Argenti P., Azzaroli, A., Caloi L., Capasso Barbato L., Di Stefano G., Esu D., Ficcarelli G., Girotti O., Kotsakis T., Masini F., Mazza P., Mezzabotta C., Palombo M.R., Petronio C., Rook L., Sala B., Sardella R., Zanalda E. and Torre D. (1997). Biochronology of selected Mammals, Molluscs, and Ostracods from the Middle Pliocene to the Late Pleistocene in Italy: the state of the art. Rivista Italiana di Paleontologia e Stratigrafia 103(3): 369-388.
Gorjanovich-Kramberger D. (1913). Fossilni rinocerotidi Hrvatske i Slavonije, s osobitim obzirom na Rhinoceros mercki iz Krapine (De rhinocerotidibus fossilibus Croatie et Slavoniae, praecipua ratione habita Rhinocerotis mercki var. Krapinensis mihi). Djela Jugoslavenske Akademije (JAZU), 22 (8): 1-70 [in Slovenian].
Groiss J.Th., Guenther A. & Keupp H. (1981). Eine quartäre Spaltenfüllung im Steinbruchgebiet Wintershof bei Eichstätt. A. Paläontologische Untersuchungen (C. antiquitatis u. D. kirchbergensis), B. Zur Grabung, C. Zur geologischen situation. Geologische Blätter für Nordost-Bayern und Angrenzende Gebiete, 31 (1-4): 165-188.
Gromova V.I. (1932). Novye materialy po chetvertichnoy faune Povolzh’ya i istorii mlekopitayushchikh Vostochnoy Evropy voobshche [New material on the Quaternary Fauna and history of Mammalia from Eastern Europe]. Trudy Komissii po Izucheniyu Chetvertichnogo Perioda, 2: 69-184 [in Russian].
Gromova V.I. (1935). Ob ostatkakh nosoroga Merka (Rhinoceros mercki Jaeg.) s Nizhney Volgi. Ueber die Reste des Merckschen Nashorn (Rhinoceros mercki Jaeg.) von der unteren Wolga. Trudy Paleontologicheskogo Instituta AN SSSR, IV: 91-131 [in Russian, German summary].
Guérin C. (1980). Les Rhinocéros (Mammalia, Perissodactyla) du Miocène Terminal au Pléistocène Supérieur en Europe Occidentale - Comparaison avec les espèces actuelles. Thèse Doctorat d’Etat et Sciences, Univ. Lyon I, Documents des Laboratoires de Géologie, 79 (1-2-3): 1-1185 [English, German, and Russian summ.].
Handa, N. and Pandolfi, L. (2016). Reassessment of the Middle Pleistocene Japanese rhinoceroses (Mammalia, Rhinocerotidae) and paleobiogeographic implications. Paleont. Res. 20: 247-260. [re-identification of Dicerorhinus nipponicus (Shikama et al. 1967) as Stephanorhinus kirchbergensis]
Hashemi, N., Ashouri, A., Aliabadian, M., Gharaie, M. H. M., Marco, A. S. and Louys, J. (2016). First report of Quaternary Mammals from the Qalehjough area, Lut desert, Eastern Iran. Palaeontologia Electronica 19(3): 1-12.
Hinton M.A.C. & Kennard A.S. (1900). Contributions to the Pleistocene geology of the Thames Valley I - The Grays Thurrock area, part 1. Essex Naturalist, 11: 336-370.
Hooijer D.A. (1961). The fossil vertebrates of Ksar’ Akil, a palaeolithic rock shelter in the Lebanon. Zoologische Verhandlungen 49: 3-68.
Hrynowiecka, Anna et al. (2020). Palaeoenvironment of the extinct rhino (Stephanorhinus kirchbergensis; Jäger, 1839) from Gorzów Wielkopolski (W Poland), pp. 57-58. In: Sobczyk A., Ratajczak-Skrzatek U., Kasprzak M., Kotowski A., Marciszak A., Stefaniak K. (eds.). Proceedings of INQUA SEQS 2020 Conference, Wrocław, Poland. University of Wrocław & Polish Geological Society. 124 pp.
Jánossy D. (1986). Pleistocene Vertebrate Faunas of Hungary. Akadémiaio Kiadó, 209 pp., Budapest [revised translation of: JÁNOSSY D., 1979 - A Magyarországi Pleisztocén Tagolása Gerinces Faunák Alapján, Akadémiaio Kiadó, Budapest].
Kahlke H.-D. (1975). Die Rhinocerotiden-Reste aus den Travertinen von Weimar-Ehringsdorf. Paläontologische Abhandlungen, A 23: 337-397.
Kahlke H.-D. (1977). Die Rhinocerotiden-Reste aus den Travertinen von Taubach bei Weimar. Quartärpaläontologie, 2: 305-359.
Kahlke H.-D. (1978). Die Rhinocerotiden-Reste aus den Travertinen von Burgtonna in Thüringen. Quartärpaläontologie, 3: 129-135.
Kahlke H.-D. (1984). Die Rhinocerotiden-Reste aus den Travertinen von Weimar (Travertine im Stadtgebiet). Quartärpaläontologie, 5: 381-387.
Kahlke, Ralf-Dietrich and Kaiser, Thomas M. (2011). Generalism as a subsistence strategy: advantages and limitations of the highly flexible feeding traits of Pleistocene Stephanorhinus hundsheimensis (Rhinocerotidae, Mammalia). Quaternary Science Reviews 30(17-18): 2250-2261. [Abstract]
Khisarova G.D. (1963). Iskopaemye kosti mlekopitayushchikh iz Koshkurganskogo grifona, Yuzhny Kazakhstan [Mammal osteological fossil remains from the Koshkurgan locality, Southern Kazakhstan]. Materialy po Istorii Fauny i Flory Kazakhstana, 4: 42-75, AN KazSSR, Izd-vo “Nauka” KazSSR, Alma-Ata [in Russian].
Kirillova, Irina V. et al. (2017). Discovery of the skull of Stephanorhinus kirchbergensis (Jäger, 1839) above the Arctic Circle. Quaternary Research 88(3): 537-550. [Abstract]
Kirillova, Irina V. et al. (2021). On Time and Environment of Stephanorhinus kirchbergensis Jäger 1839 (Mammalia, Rhinoceratidae) in Altai and Northeastern Russia. Biology Bulletin 48: 1674-1687. https://doi.org/10.1134/S1062359021090077
Klein, R.G., Scott, K., 1986. Re-analysis of faunal assemblages from the Haua Fteah and other late Quaternary archaeological sites in Cyernaican Libya. J. Archaeol. Sci. 13, 515–542.
Kosintsev, Pavel A. et al. (2022). Mammalian fauna of the Late Pleistocene from the Barsuchiy Dol Cave (Southern Urals). Russian J. Theriol. 21(2): 180-191.
Kowalski K. (1959). Katalog Ssaków Plejstocenu Polski [Catalogue of the Pleistocene Mammals of Poland]. Polska Akademia Nauk (Institut Zoologii), Oddz. Kraków: 153-154 [in Polish].
Kozhamkulova B.S. & Kostenko N.N. (1984). Vymershie Zhivotnye Kazakhstana - Paleogeografya Poznego Kaynozoya [Extinct animals of Kazakhstan - The paleogeography of the Late Cenozoic]. AN KazSSR, 103 pp., Izd-vo Nauka KazSSR, AlmaAta [in Russian].
Krakhmalnaya, T. V., Kovalchuk, O. M. and Derkach, T. G. (2018). Rhinoceroses from the paleontological collection of the Zaporozhye Regional Museum of Local History (Ukraine): Systematic and Ecological Diversity. International Symposium "Functional Ecology of Animals".
Kurtén B. (1968). Pleistocene Mammals of Europe. Weidenfeld & Nicolson ed., 317 pp., London.
Lacombat F. (2005). Les rhinocéros fossiles des sites préhistoriques de l’Europe Méditerranéenne et du Massif Central - Paléontologie et implications biochronologiques. British Archaeological Reports, 1419: 1-175.
Lacombat, F. 2006. Morphological and biometrical differentiation of the teeth from Pleistocene species of Stephanorhinus (Mammalia, Perissodactyla, Rhinocerotidae) in Mediterranean Europe and the Massif Central, France. Palaeontographica Abteilung, 274:71-111.
Lartet E. (1867). Notes sur deux têtes de Carnassiers fossiles (Ursus et Felis) et sur quelques débris de Rhinocéros provenant des découvertes faites par M. Bourguignat dans les cavernes du Midi de la France. Annales des Sciences Naturales (Zoologie), 5 (8): 157-194.
Lee Yung-Jo (2001). Bone Remains from Sae-kul and Cheonyo-kul of the Turubong Cave Complex in Korea. Proceedings of the 3rd International Scientific Conference “Asia and the Middle Pleistocene in Global Perspective”, 3: 14-17.
Lobachev, Y. V., Shpansky, А.V., Bondarev, A.A., Lobachev, A.Y., Vasiliev, S.K., Klementev, A.M., Grebnev, I.E., and Silaev, V.I. (2021). New findings of Stephanorhinus kirchbergensis in Siberia. Palaeontologia Electronica 24(1): a14. https://doi.org/10.26879/734
Loose, H. K. (1975). Pleistocene Rhinocerotidae of Western Europe with Reference to the Recent Two-Horned Species of Africa and S.E. Asia. Scripta Geologica 33: 1-59.
Lubicz-Niezabitowski K.E. (1926). Szczatki Nosorozca Wlochatego (Rhinoceros antiquitatis Blum.) - Znalezione na Ziemi Wielkopolskiej [Les Débris du Rhinoceros antiquitatis Blum. - Trauvés sur les Territoires de la Grande Pologne] [in Polish]. Rocznik Muzeum Wielkopolski, 2: 1-52.
Malez M. (1970). Rezultati revizija pleistocenska fauna iz Krapine [Results of the revision of the Pleistocene fauna from Krapina]. In Malez M. (ed.), “Krapina 1899-1969”, pp. 45-56 [in Croatian, German, and English].
Malez M. (1986). Die quartären Vertebraten-Faunen in der SFR Jugoslawien. Quartärpaläontologie, 6: 101-117 [English and Russian summ.].
Markova A.K., Puzachenko A.Y., van Kolfschoten T. 2010 The North Eurasian mammal assemblages during the end of MIS 3 (Brianskian–Late Karginian–Denekamp Interstadial). Quaternary International 212(2), 149-158.
Mayer G. (1971). Beiträge zur Geschichte der Badischen Landessammlungen für Naturkunde in Karlsruhe. Der Schädel des Dicerorhinus mercki (kirchbergensis) (Jäger) var. Brachycephalus Schroeder von Daxlanden und seine Geschichte. Beiträge Naturkunde Forschungen Südwest Deutschland, 30 (2): 157-163.
Meyer H.v. (1863-64). Die diluvianen Rhinoceros-Arten. Palaeontographica, 11: 233-283.
Mirosław-Grabowska, Joanna et al. (2020). Eemian and Vistulian palaeolake horizons in Gorzów Wielkopolski (NW Poland) on the basis of isotopic and geochemical data, pp. 87-88. In: Sobczyk A., Ratajczak-Skrzatek U., Kasprzak M., Kotowski A., Marciszak A., Stefaniak K. (eds.). Proceedings of INQUA SEQS 2020 Conference, Wrocław, Poland. University of Wrocław & Polish Geological Society. 124 pp.
Morris J. (1836). On a freshwater deposit containing mammalian remains, recently discovered at Grays, Essex. Magazine of Natural History, s. 1, 9: 261-264.
Pandolfi, Luca. (2021). Understanding the variability of Early Pleistocene Stephanorhinus (Mammalia, Rhinocerotidae): implications for taxonomy and palaeobiogeography, pp. 84-85. In: Ratajczak-Skrzatek U., Kovalchuk O., Stefaniak K. (eds.). Proceedings of INQUA SEQS 2021 Conference, Wrocław, Poland. University of Wrocław & Polish Geological Society. 119 pp.
Pandolfi, Luca. (2023). Reassessing the phylogeny of Quaternary Eurasian Rhinocerotidae. Journal of Quaternary Science. https://doi.org/10.1002/jqs.3496
Pandolfi, Luca and Marra, Fabrizio. (In Press, 2015). Rhinocerotidae (Mammalia, Perissodactyla) from the chrono-stratigraphically constrained Pleistocene deposits of the urban area of Rome (Central Italy). Geobios. doi:10.1016/j.geobios.2015.02.003 [Abstract]
Pandolfi, Luca and Petronio, Carmelo. (2011). The small-sized rhinoceroses from the Late Pleistocene of Apulia (southern Italy). Rivista Italiana di Paleontologia e Stratigrafia 117(3): 509-520.
Persico, D., Billia, E. M. E., Ravara, S. and Sala, B. (2015). The skull of Stephanorhinus kirchbergensis (Jäger, 1839) (Mammalia, Rhinocerotidae) from Spinadesco (Cremona, Lombardia, Northern Italy): morphological analyses and taxonomical remarks – an opportunity for revising the three other skulls from the Po Valley. Quaternary Science Reviews 109: 28-37. [Abstract]
Pushkina D. 2007 The Pleistocene easternmost distribution in Eurasia of the species associated with the Eemian Palaeoloxodon antiquus assemblage. Mammal Review 37(3), 224-245.
Radulescu C., Samson P-M., Mihaila N. & Kovacs A. (1965). Contributions à la connaissance des faunes de Mammifères pléistocènes de la Dépression de Brasov (Roumanie). Eiszeitalter und Gegenwart, 16: 132-188.
Rakovec I. (1933). Coelodonta mercki Jäger iz Dolarjeve jame pri Logatcu [Coelodonta mercki Jäger aus der Höhle “Dolarjeva jama” bei Logatec]. Prirodoslov Razprave [Dissertation on Natural History], SAZU (Academia Scientiarum et Artes Slovenica), 2: 5-41 [in Slovenian].
Rakovec I. (1958). Pleistocenski sesalci iz jame pri Crnem Kalu v Istri [The Pleistocene mammalia from the Crni Kal cave (Northern Istria)]. Prirodoslov. Razprave [Dissertation on Natural History], Academia Scientiarum et Artium Slovenica (Slovenska Akademija Znanosti i Umetnosti [SAZU]), cl. 4, 4: 367-433 [in Slovenian].
Salov I.N. (1957). Nakhodki iskopaemoy chetvertichnoy fauny v Smolenskoy oblasti [Finds of fossil Quaternary fauna in the Smolensk region]. Smolensky Oblastnoy Kraevedchesky Muzey, 2: 311-333 [in Russian].
Samson P. & Nadisan I. (1970). Sur la présence de Dicerorhinus kirchbergensis (Jäger) dans les travertins du Bassin de Borsec (Département de Harghita). Travaux de l’Institute “E. Racovita”, 9: 245-251.
Schroeder H. (1903). Die Wirbelthier-Fauna des Mosbacher Sandes - Gattung Rhinoceros.Abhandlungen der Kaiserlich-Königlichen Preussichen Geologischen Landesanstalt, NF 18: 1-143.
Scott, K. (2001). Late Middle Pleistocene Mammoths and Elephants of the Thames Valley, Oxfordshire, pp. 247-254. In: Cavarretta, C., Gioia, P., Mussi, M. and Palombo, M. R. (eds.). The World of Elephants – International Congress, Rome. Proceedings of the 1st international congress – Consiglio nazionale delle ricerche, Rome. [automatic download]
Shikama, T., Hasegawa, Y. and Okafuji, G. (1967). On a rhinocerotid skull from Isa (Yamaguchi Prefecture, Japan). Bull. Nat. Sci. Mus. (Tokyo) 10: 455-462.
Shpansky, A. V. and Boeskorov, G. G. (2018). Northernmost Record of the Merck’s Rhinoceros Stephanorhinus kirchbergensis (Jäger) and Taxonomic Status of Coelodonta jacuticus Russanov (Mammalia, Rhinocerotidae). Paleontological Journal 52(4): 445-462.
Simionescu I. (1939-40). Rhinoceros mercki in Rumänien. Buletin Stiintific (Sectiunea de Stiinte Biologice, Agronomice, Geologice si Geografice), 22 (10): 429-432.
Smith F.A., Lyons S.K., Ernest S.K.M., Jones K.E., Kaufman D.M., Dayan T., Marquet P.A., Brown J.H., Haskell J.P. 2003 Body mass of late Quaternary mammals. Ecology 84(12), 3403-3403.
Stäesche K. (1941). Nashörner der Gattung Dicerorhinus aus Diluvium dem Württembergs. Abhandlungen der Reichsanstalt Bodenforschung, 200: 1-148.
Stefaniak, Krzysztof et al. (In Press, 2020). Browsers, grazers or mix-feeders? Study of the diet of extinct Pleistocene Eurasian forest rhinoceros Stephanorhinus kirchbergensis (Jäger, 1839) and woolly rhinoceros Coelodonta antiquitatis (Blumenbach, 1799). Quaternary International. https://doi.org/10.1016/j.quaint.2020.08.039 [Abstract]
Stewart J.R. 2007 Neanderthal extinction as part of the faunal change in Europe during Oxygen Isotope Stage 3. Acta Zoologica Cracoviensia - Series A: Vertebrata 50(1-2), 93-124.
Strizheva I.M. (1991). Kostnye ostatki krupnykh mlekopitayushchikh v rayone Samarskoy Luki [Osteological remains of large mammals from the Samara Luka district]. Samarskaya Luka, 2: 197-205 [in Russian].
Sun, Boyang et al. (2022). New material of Stephanorhinus (Rhinocerotidae, Mammalia) from Jinyuan Cave, Luotuo Hill in Dalian, Northeast China. Historical Biology. https://doi.org/10.1080/08912963.2022.2130793
Teilhard de Chardin P. & Pei W.C. (1941). The fossil Mammals of locality 13 in Choukoutien. Palaeontologia Sinica, s. C, 11(4): 1-105.
Thenius E. (1956-59). Die jüngpleistozäne Wirbeltierfauna von Willendorf i. d. Wachau, N.Ö. Mitteilungen der Prähistorischen Kommission Österr. Akademie der Wissenschaften, 8/9: 133-170.
Tong H., Wu X. 2010 Stephanorhinus kirchbergensis (Rhinocerotidae, Mammalia) from the Rhino Cave in Shennongjia, Hubei. Chinese Science Bulletin 55(12), 1157-1168.
Toula, F. (1902). Das Nashorn von Hundsheim Rhinoceros (Ceratorhinus Osborn) hundsheimensis nov. form. mit Ausführungen über die Verhältnisse von elf Schädeln von Rhinoceros (Ceratorhinus) sumatrensis. Abhandlungen der Kaiserlich-Königlichen Geologischen Reichsanstalt, 19 (1): 1-92.
Toula F. (1907). Rhinoceros mercki Jäger in Österreich. Jahrbuch der Kaiserlich-Königlichen Geologischen Reichsanstalt, 57 (3): 445-454.
van Asperen, Eline N. and Kahlke, Ralf-Dietrich. (2015). Dietary variation and overlap in Central and Northwest European Stephanorhinus kirchbergensis and S. hemitoechus (Rhinocerotidae, Mammalia) influenced by habitat diversity: “You'll have to take pot luck!” (proverb). Quaternary Science Reviews 107: 47-61. [Abstract]
Wang T. (1961). [The occurence of Stegodon and Merck’s Rhinoceros near Taiyuan, Shansi]. Vertebrata PalAsiatica, 6(2): 160-162 [in Chinese].
Welker F, Smith GM, Hutson JM, Kindler L, Garcia-Moreno A, Villaluenga A, Turner E, Gaudzinski-Windheuser S. (2017) Middle Pleistocene protein sequences from the rhinoceros genus Stephanorhinus and the phylogeny of extant and extinct Middle/Late Pleistocene Rhinocerotidae. PeerJ 5: e3033.
West R.G., Lambert C.A. & Sparks B.W. (1964). Interglacial deposits at Ilford (Essex). Philosophical Transactions of the Royal Society of London, s. B, 738, 247: 185-212.
Wüst E. (1909). Das Vorkommen von Rhinoceros merckii Jäger in den oberen Travertinen von Ehringsdorf bei Weimar und seine Bedeutung für die Beurteilung der Klimaschwankungen das Eiszeitalters. Centralblatter für Mineralogie, Geologie und Paläontologie, 3: 23-25.
Wüst E. (1911). Zwei bemerkenswerte Rhinoceros-Schädel aus dem Pliozän Thüringens. Palaeontographica, 58: 133-138.
Wüst E. (1914). Rhinoceroszähne aus dem Sande von Mauer bei Heidelberg. In Freudenberg W. (ed.), Die Säugetiere des Älteren Quartärs von Mitteleuropa mit Besonder Berücksichtigung der Fauna von Hundsheim und Deutschaltenburg in Nieder- Österreich. Geologische und Paläontologische Abhandlungen, NF 12 (4-5): 23-26 + 475-478.
Wüst, E. (1922). Beitraege zur Kenntnis der diluvianen Nashoerner Europas. Centralblatter für Mineralogie, Geologie und Paläontologie, 20: 641-656; 21: 680-688.
Xu X. (1986). [Dicerorhinus kirchbergensis (Jäger, 1839) from the Late Middle Pleistocene mammalian fauna of Anping (Liaoning)]. Vertebrata PalAsiatica, 24 (3): 229-241 [in Chinese].
Zeuner, F. E. (1945). New reconstruction of the woolly Rhinoceros and Merck’s Rhinoceros. Proceedings of the Linnaean Society of London 156(3): 1-58.
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