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Diprotodon optatum Owen, 1838a:362-363, xix

Giant wombat, ?yamuti (Tunbridge, 1988, 1991:72,89)

 

 

Taxonomy & Nomenclature

Synonyms: Dinotherium australe Owen, 1843; Diprotodon australis Owen, 1844; Diprotodon annextans McCoy, 1861; Diprotodon minor Huxley, 1862; Diprotodon longiceps McCoy, 1865; Diprotodon loderi Krefft, 1873a; Diprotodon bennetti Krefft, 1873b; Diprotodon bennettii Owen, 1877

 

Conservation Status

Extinct

Last record: 44ka (Johnson et al., 2016)

 

Diprotodon was the first Australian megafauna species to be described, based upon fossils collected by the explorer Thomas Mitchell in 1830 from the world famous Wellington Caves. It also appears to have been one of the most recent megafauna extinctions (Johnson et al. 2016), along with Zygomaturus trilobus and perhaps some of the sthenurine kangaroos.

 

Distribution

Australia

 

Anatomy & Morphology

A mass of 1000kg (Johnson & Prideaux, 2004:557) and then 2700kg has been given (Johnson, 2006:18). The maximum weight of males of the Giant wombat (D. optatum) have been estimated to be 2786kg (Wroe et. al. 2004), which would make it the largest known marsupial to have ever lived.

 

Biology & Ecology

It was a browser (Johnson, 2006:18).

 

Hypodigm

Holotype: BM M10796 (Dawson, 1985:66)

Type locality: "coll. by Mitchell, 1830, in "the large cave"" (Dawson, 1985:66)

Other specimens:

QMF44649 ("P3")
NMV P31299 (cranium) (Sharp, 2014)
NMV P151802 (lower mandible) (Sharp, 2014)
NMV P157382 (lower mandible) (Sharp, 2014)

 

Media

"Diprotodon teeth excavated from Wellington Caves. Australian Museum."

Source: MacLennan, Sally. (2020). At the Museum: Have you even seen a two tonne wombat? Central Western Daily, available from: https://www.centralwesterndaily.com.au/story/6837198/at-the-museum-have-you-even-seen-a-two-tonne-wombat/

 

Agnozoology

A traditional bunyip account is that of an encounter by mineralogist Charles Bailly with an unknown animal bellowing in the reeds up the Swan River, Western Australia, in June 1801. Although (Michell & Rickard, 1982) hypothesise that it might refer to living Diprotodon.

The explorer Hamilton Hume encountered a 'hippopotamus' in Lake Bathurst, New South Wales in 1821, which might refer to a living Giant wombat (Gilroy, 1976; Michell & Rickard, 1982). Michell & Rickard (1982) included this and another possible encounter.

 

References

Original scientific description:

Owen, Richard. (1838a). Letter in: Mitchell, Thomas L. Three Expeditions into the Interior of Eastern Australia, with descriptions of the recently explored region of Australia Felix, and of the present colony of New South Wales. Vol. 1. London: T. & W. Boone, 343 pp. [relevant citation?]

 

Other references:

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Anderson, C. and Fletcher, H. O. (1934). The Cuddie Springs bone bed. The Australian Museum Magazine 5(5): 152-158.

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Scott, H. H. and Lord, C. E. (1920b). Studies in Tasmanian mammals, living and extinct. Number II. Section 1: The history of the genus Nototherium. Section 2: The osteology of the cervical vertebrae of Nototherium mitchelli. Papers and Proceedings of the Royal Society of Tasmania 1920: 17-32.

Sharp, Alana Clare. (2014). Three dimensional digital reconstruction of the jaw adductor musculature of the extinct marsupial giant Diprotodon optatum. PeerJ 2: e514.

Sharp, Alana Clare. (2015). Cranial form and function of the largest ever marsupial, Diprotodon optatum Owen, 1838 (Marsupialia: Diprotodontinae). Doctorate thesis, Monash University. [Abstract]

Sharp, Alana Clare and Rich, T. H. (2016). Cranial biomechanics, bite force and function of the endocranial sinuses in Diprotodon optatum, the largest known marsupial. Journal of Anatomy. doi: 10.1111/joa.12456 [Abstract]

Smith F.A., Lyons S.K., Ernest S.K.M., Jones K.E., Kaufman D.M., Dayan T., Marquet P.A., Brown J.H., Haskell J.P. 2003 Body mass of late Quaternary mammals. Ecology 84(12), 3403-3403.

Stephenson, N. G. (1963). Growth gradients among fossil monotremes and marsupials. Palaeontology 6(4): 615-624.

Stirling, E. C. (1893). [Extract from a letter concerning the discovery of Diprotodon and other mammalian remains in South Australia]. Proceedings of the Zoological Society, London 1893: 473-475.

Stirling, E. C. (1901). [Diprotodon australis.) Museum Journal. (London) 1: 114-115 (1901-1902)

Stirling, E. C. (1907). Reconstruction of Diprotodon from the Callabonna deposits, South Australia. Nature (London) 76: 543-544, figs. 1, 2.

Stirling, E. C. and Zietz, A. H. C. (1899). Fossil remains of Lake Callabonna. Part I. Description of the manus and pes of Diprotodon australis, Owen. Memoirs of the Royal Society of South Australia 1: 1-40, pis. I-XVIII. Abstr., Nature (London) 61: 275-278, 2 figs. (1900). [First page]

Tedford, Richard H. (1973). The Diprotodons of Lake Callabonna. Australian Natural History 17(11): 349-354.

Tedford, Richard H. (1994). Lake Callabonna: 'Veritable necropolis of gigantic extinct marsupials and birds'. Abstracts of the fourth conference on Australian vertebrate evolution, palaeontology and systematics, Adelaide, 19-21 April, 1993. Records of the South Australian Museum 1994. [Abstract] [curiously: "possibly a second, smaller species [of Diprotodon]"]

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Tedford, R.H., Wells, R.T., and Barghoorn, S.F. (1992). Tirari Formation and contained faunas, Pliocene of the Lake Eyre Basin, South Australia. The Beagle, Records of the Northern Territory Museum of Arts and Sciences 9: 173-194.

Tindale, N. B., Fenner, F. J. and Hall, F. J. (1935). Mammal bone beds of probable Pleistocene age, Rocky River, Kangaroo Island. Transactions of the Royal Society of South Australia 59: 103-106.

Travouillon, Kenny J., Jackson, Stephen, Beck, Robin M. D., Louys, Julien, Cramb, Jonathan, Gillespie, Anna, Black, Karen, Hand, Suzanne, Archer, Michael, Kear, Benjamin, Hocknull, Scott, Phillips, Matthew, McDowell, Matthew, Fitzgerald, Erich M. G., Brewer, Phillipa and Price, Gilbert J. (2024). Checklist of the Fossil Mammal Species of Australia and New Guinea.  Available from: https://www.australasianpalaeontologists.com/national-fossil-species-lists [Accessed 24 November 2024]

Trezise, P. (1993). Dream road: A journey of discovery. Sydney: Allen and Unwin. ["[proposes] that an image from Cape York Peninsula is of Diprotodon"*]

Trueman, Clive N. G., Field, Judith H., Dortch, Joe, Charles, Bethan and Wroe, Stephen. (2005). Prolonged coexistence of humans and megafaunain Pleistocene Australia. PNAS 102(23): 8381-8385. https://doi.org/10.1073/pnas.0408975102

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Zietz, A. H. C. (1890b). (Abstract of Proceedings). Transactions of the Royal Society of South Australia 13: 245. [relevant citation?]

http://museum.wa.gov.au/sites/default/files/VOL.1%20PART%201%201910%20P509.941%20REC.pdf

*This quote is taken from: Bednarik, Robert G. (2013). Myths About Rock Art. Journal of Literature and Art Studies 3(8): 482-500.

http://www.abc.net.au/news/2016-11-24/teeth-of--rhinoceros-sized-wombat-found-at-lancefield-swamp/8053196

https://twilightbeasts.wordpress.com/2016/10/12/squishy-bear-face/

https://twilightbeasts.wordpress.com/2014/10/06/an-adorable-goofy-looking-giant/

https://www.australianmuseum.net.au/blogpost/museullaneous/meet-darren-the-diprotodon

https://www.biodiversitylibrary.org/item/84659#page/253/mode/1up

http://extinctanimals.proboards.com/thread/16034/diprotodon-optatum

https://www.abc.net.au/news/2019-06-28/giant-wombat-monaro-fossil-museum/11258792

 

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