Raphus cucullatus Linnaeus, 1758:155




Taxonomy & Nomenclature

Synonym/s: Struthio cucullatus Linnaeus, 1758:155; Didus ineptus Linnaeus, 1766; Cygnus cucullatus Nieremberg, 1635:231; Didus nazarenus Bartlett, 1851:284?


Conservation Status


Last record: c.1638 (Fuller, 2002)

IUCN status: Extinct


The Dodo is probably the most famous of all recently extinct species, having entered the vocabulary of ordinary people who have no particular knowledge of the animal ("dead as a dodo", "to go the way of the dodo" etc.). It was driven extinct within roughly 50 years of its initial contact with Europeans (Fuller, 2002). Reports of its persistence occur to the present (Shuker, 1989,1998,1999:47).



Mauritius, Mascarene Islands


Biology & Ecology




The Grant Museum of Zoology:

Y86 (casts of head and foot)

Y105 (partial skeleton)


Other specimens:

NMS.Z.1993.13 (Brassey et al. 2016)

NHMUK A.9040 and S/1988.50.1 (Brassey et al. 2016)






Original scientific description:

Linnaeus, Carolus. (1758). Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis. Stockholm, Sweden.


Other references:

Abrahamson, W. G. (1989). Plant-animal interactions: An overview. In: Abrahamson, W. G. (ed.), Plant-animal interactions. McGraw-Hill, NY, pp. 1-20.

Ali, S. (1968). Dodo in Maalvi and Rahman A. (eds.). Jahangir the Naturalist. new Delhi: National Institute of Sciences of India.

Angst, Delphine. (2018). Les dodos : des oiseaux toujours très mystérieux. Géochronique 146: 63-71.

Angst, Delphine, Buffetaut, E. and Abourachid, A. (2011a). The end of the fat dodo? A new mass estimate for Raphus cucullatus. Naturwissenschaften 98(3): 233-236. [Abstract]

Angst, Delphine, Buffetaut, E. and Abourachid, A. (2011b). In defence of the slim dodo: a reply to Louchart and Mourer-Chauviré. Naturwissenschaften 98(4): 359-360.

Angst, Delphine, Chinsamy, A., Steel, L. and Hume, Julian Pender. (2017). Bone histology sheds new light on the ecology of the dodo ([i]Raphus cucullatus[/i], Aves, Columbiformes). Scientific Reports 7(1): 7993.

Anonymous, 1868: Zpráva o Museu království Českého od 1. září do konce listopadu 1868 [Report on the Museum of the Kingdom of Bohemia from 1 September to the end of November 1868]. Časopis Českého Musea 42: 406-409. [In Czech.]

A True Report. (1599). A True Report of the gainefull, prosperous and speedy voyage to Java in the East Indies, performed by a fleete of eight ships of Amsterdam: which set forth from texell in Holland, the first of Maie 1598 Stilo Neue. Whereof foure returned againe the 19. july Anno 1599. In less than 15. Months, the other foure went forward from Java for the Moluccas. London.

Baker, R. A. and Bayliss, R. A. (2002). Alexander Gordon Melville (1819-1901): the Dodo, Raphus cucullatus (L., 1758) and the genesis of a book. Archives of Natural History 29(1): 109-118. [Abstract]

Bartlett, A. D. (1851). On some Bones of Didus. Proceedings of the Zoological Society of London 19: 280-284.

Bartlett, A. D. (1854). On some Bones of Didus. Ann. Mag. Nat. Hist. (2) XIV: 297-301.

Begon, M., Harper, J. L. and Townsend, C. R. 1990. Ecology: Individuals, populations and communities. Blackwell, Oxford.

Bergman, Jerry. (2005). The History of the Dodo Bird and the Cause of Its Extinction. Perspectives on Science and Christian Faith 57(3): 221-229.

BirdLife International. (2012). Raphus cucullatus. In: IUCN 2012. IUCN Red List of Threatened Species. Version 2012.1. ( Downloaded on 01 October 2012.

BirdLife International. (2016). Raphus cucullatus. The IUCN Red List of Threatened Species 2016: e.T22690059A93259513. Downloaded on 12 July 2020.

Blainville, H. D. (1829). Memoire sur le Dodo. Nouv. Ann. du Musee d'Hist. Nat.4: 1-46.

Blainville, H. D. (1835). Memoire sur le Dodo, autrement. Nouv. Ann. du Mus. 4: 1-36.

Brassey, C. A., O’Mahoney, T. G., Kitchener, A. C., Manning, P. L. and Sellers, W. I. (2016). Convex-hull mass estimates of the dodo (Raphus cucullatus): application of a CT-based mass estimation technique. PeerJ 4: e1432.

Broderip, W. J. (1837). Dodo. Penny Cyclopedia 9: 47.

Brom, T. G. and Prins, T. G. (1988). A skeleton of the dodo Raphus cucullatus (L.) new to ornithology. Bijdragen tot de Dierkunde 58: 155-158.

Brom, T. G. and Prins, T. G. (1989). Microscopic investigation of feather remains from the head of the Oxford dodo, Raphus cucullatus. Journal of Zoology (London) 218(2): 233-246.

Brooks, T. 2000. Extinct species. In: BirdLife International (ed.), Threatened Birds of the World, pp. 701-708. Lynx Edicions and BirdLife International, Barcelona and Cambridge, U.K.

Brown, Clare M. (2020). Harry Pasley Higginson and his role in the re-discovery of the dodo (Raphus cucullatus). Archives of Natural History 47(2): 381-391. [Abstract]

Catling, Paul M. (2001). Extinction and the importance of history and dependence in conservation. Biodiversity 2(3): 2-14. [Abstract]

Cauche, F. (1651). Relation du Voyage de F. Clauche, in relations veritables et curieuses de I'Isle de Madagascar. London.

Cheke, Anthony S. (1985). Dodo. Pp.152 in Campbell, Bruce & Lack, Elizabeth (eds.). 1985. A dictionary of birds. Calton, Staffordshire, UK: T. & A.D. Poyser. 670pp.

Cheke, Anthony S. (1987). An ecological history of the Mascarene Islands, with particular reference to extinctions and introductions of land vertebrates, pp. 5-89. In: Diamond, A. W. (ed.). Studies of Mascarene Island birds. Cambridge: Cambridge Univ. Press.

Cheke, Anthony S. (2006). Establishing extinction dates - the curious case of the Dodo Raphus cucullatus and the Red Hen Aphanapteryx bonasis. Ibis 148: 155-158.

Cheke, Anthony S. (2010). Is the bird a Dodo? The wildlife of a mid-seventeenth century drawing of Dutch Mauritius. Archives of Natural History 28(3): 347-351.

Cheke, Anthony S. (2013). Extinct birds of the Mascarenes and Seychelles - a review of the causes of extinction in the light of an important new publication on extinct birds. Phelsuma 21: 4-19.

Cheke, Anthony S. (2015). Speculation, statistics, facts and the Dodo's extinction date. Historical Biology 27(5): 624-633. [Abstract]

Cheke, Anthony S. and Beentje, Henk. (2016). Citizen animals, dodo mayors – the curious account of a visit to Mauritius in 1632. Phelsuma 24: 1-5.

Cheke, Anthony S. and Beentje, Henk. (2019). Exploring At Second-Hand: Separating the Editor from the Traveller in Soeteboom’s Version of Van West-Zanen’s Sojourn in Mauritius in 1602. Journal of the Royal Society of Arts & Sciences of Mauritius 1: 37-47.

Cheke, Anthony S., Gardner, Tony, Jones, Carl G., Owadally, A.Wahab & Staub, France. 1984. Did the dodo do it ? Animal Kingdom 87(1): 4-6.

Cheke, Anthony S. and Hume, Julian Pender. (2008). Lost Land of the Dodo: An Ecological History of Mauritius, Réunion & Rodrigues. New Haven and London: Yale University Press. 464 pp.

Cheke, Anthony S. and Parish, Jolyon. (2015). The ‘Bundi dodo’: a drawing copying European animals – a curious byway in nineteenth century Indian art. Historical Biology. DOI: 10.1080/08912963.2015.1079629 [Abstract]

Cheke, Anthony S. and Parish, Jolyon. (2020). The Dodo and the Red Hen, A Saga of Extinction, Misunderstanding, and Name Transfer: A Review. Quaternary 3(1): 4.

Chubb, E. C. (1919). A skeleton of the dodo ([i]Didus ineptus[/i]). Annals of theDurban Museum 2: 97-99.

Claessens, L. P. A. M., and J. P. Hume. 2015. Provenance and history of the Thirioux dodos; pp. 21–28 in L. P. A. M. Claessens, H. J. M. Meijer, J. P. Hume, and K. F. Rijsdijk (eds.), Anatomy of the Dodo (Raphus cucullatus L., 1758): An Osteological Study of the Thirioux Specimens. Society of Vertebrate Paleontology Memoir 15. Journal of Vertebrate Paleontology 35(6, Supplement). [Abstract]

Claessens, L. P. A. M., H. J. M. Meijer, and J. P. Hume. 2015. The morphology of the Thirioux dodos; pp. 29–187 in L. P. A. M. Claessens, H. J. M. Meijer, J. P. Hume, and K. F. Rijsdijk (eds.), Anatomy of the Dodo (Raphus cucullatus L., 1758): An Osteological Study of the Thirioux Specimens. Society of Vertebrate Paleontology Memoir 15. Journal of Vertebrate Paleontology 35(6, Supplement). [Abstract]

Claessens, L. P. A. M., H. J. M. Meijer, J. P. Hume and Kenneth F. Rijsdijk. (2015). Preface, pp. 1-2 in L. P. A. M. Claessens, H. J. M. Meijer, J. P. Hume, and K. F. Rijsdijk (eds.), Anatomy of the Dodo (Raphus cucullatus L., 1758): An Osteological Study of the Thirioux Specimens. Society of Vertebrate Paleontology Memoir 15. Journal of Vertebrate Paleontology 35(6, Supplement).

Clark, George. (1866). Account of the late discovery of dodos' remains in the island of Mauritius. Ibis 8(2): 141-146. [1865?]

Clusius, C. (1605). Exoticorum libri decem. Leyden.

Courtenay Latimer, Marjorie. (1953). A Dodo Egg (Letters to the editors). South African Journal of Science 49(6): 208-210.

Crawley, M. J. 1983. Herbivory: The dynamics of animal-plant interactions. - Univ. California Press, Berkeley, CA.

de Boer, Erik J. et al. (2015). A deadly cocktail: How a drought around 4200 cal. yr BP caused mass mortality events at the infamous ‘dodo swamp’ in Mauritius. The Holocene 25(5): 758-771. [Abstract]

Den Hengst, Jan. (2003). The Dodo: the bird that drew the short straw. Marum: Art Revisited, 119 pp.

Doherty, Berlie and Doherty, Janna. (1989). Tilly Mint And The Dodo. London: HarperCollins Publishers.

Duncan, J. S. (1828). A summary review of the authorities on which natu-ralists are justified in believing that the dodo, [i]Didus ineptus[/i] Linn.,was a bird existing in the Isle of France, or the neighbouring islands until a recent period. Zoological Journal 3: 554-567.

Durrell, Gerald. (1977). Golden bats and pink pigeons: A journey to the flora and fauna of a unique island. New York: Simon and Schuster.

Edwards, George. (1760). Gleanings of Natural History, Part II. London: Self published.

Faaborg, J. 1988. Ornithology: An ecological approach. - Prentice-Hall, NJ.

Feinsinger, P. 1983. Coevolution and pollination. In: Futuyma, D. J. and Slatkin, M. (eds), Coevolution. Sinauer, Sunderland, MA, pp. 282-308.

Freeman, Carol J. (2007). Extinction, agency, and representation: The case of the dodo . . . and a word about the devil. In: Animals and Society II: Considering Animals, July, Hobart, Tasmania. [Summary]

Freeman, Carol J. (2011). Extinction, Representation, Agency: The Case of the Dodo, pp. 153-167. In: Freeman, Carol, Leane, Elizabeth and Watt, Yvette (eds.). Considering Animals: Contemporary Studies in Human-Animal Relations. Surrey, England and Burlington, USA: Ashgate Publishing.

Frič, A. (1861). Cesta do Londýna [A trip to London]. Živa 9: 121-144. [In Czech.]

Friedmann, Herbert. (1956). New light on the Dodo and its illustrators, pp. 475-481. In: Smithsonian Report for 1955. Washington.

Fuller, Errol. (1988). Extinct Birds. New York: Facts on File Publications. 256 pp.

Fuller, Errol. (2002). Dodo: From extinction to icon. London (UK): Collins. 192 pp.

Fulton, Graham. (2013). Dodo, Raphus cucullatus, in the Macleay Museum, The University of Sydney, Australia. Archives of Natural History 40(1): 171-174. [Abstract]

Futuyma, D. J. (1979). Evolutionary biology. Sinauer, Sunderland, MA.

van der Geer, Alexandra A. E. et al. (2021). The changing face of the dodo (Aves: Columbidae: Raphus cucullatus): iconography of the Walghvogel of Mauritius. Historical Biology 34(4): 648-657.

Geisler, M. (1994). The Dodo---an extinct species. Torgos 24: 6-7.

Gervais, Paul. (1844). Atlas de zoologie: ou collection de 100 planches. Paris: Germer Bailliere. [pl. 15]

Gilpin, M. E. and Soule, M. E. 1986. Minimum viable populations: processes of species extinction. - Sinauer, Sunderland, MA.

Gold, Maria Eugenia Leone, Bourdon, Estelle and Norell, Mark A. (2016). The first endocast of the extinct dodo ([i]Raphus cucullatus[/i]) and an anatomical comparison amongst close relatives (Aves, Columbiformes). Zoological Journal of the Linnean Society 177(4): 950-963. [Abstract]

Gosling, William D., de Kruif, Jona, Norder, Sietze J., de Boer, Erik J., Hooghiemstra, Henry, Rijsdijk, Kenneth F. and McMichael, Crystal N. H. (2017). Mauritius on fire: Tracking historical human impacts on biodiversity loss. Biotropica. DOI: 10.1111/btp.12490 [Abstract] [relevant citation?]

Gould, Stephen Jay. (1996). The dodo and the caucus race. Nat Hist. 105: 22-33.

Green, Lawrence G. (1936). Secret Africa. London: Stanley Paul. ["Natives of Mauritius told Lawrence G. Green in the 1930s that Dodos still existed in caves and mountains on remote parts of the island." (source)]

Grihault, Alan. (2005). Dodo: the bird behind the legend. Port Louis (Mauritius): Imprimerie & Papeterie Commerciale, 171 pp.

Hachisuka, M. (1953). The Dodo and kindred birds, or the extinct birds of the Mascarene Islands. London (UK): HF & G Witherby. 250 pp.

Halliday, Tim. (1978). Vanishing Birds: Their Natural History and Conservation. London: Sidgwick & Jackson.

Helfferich, C. (1990). The Turkey and the Tambalacoque Tree. Unpublished?

Hengst, Jan Den. (2009). The dodo and scientific fantasies: durable myths of a tough bird. Archives of Natural History 36(1): 136-145. [Abstract]

Herbert, Thomas. (1634). A relation of some yeares' travaile, begunne Anno 1626, into Afrique and the greater Asia, especially the territories of the Persian Monarchie, and some parts of the Oriental Indies and Isles adiacent. London: W. Stansby & J. Bloome.

Hershey, D.R. (2000). The truth behind some great plant stories. American Biology Teacher 62: 408-413.

Hershey, D. R. (2004). The widespread misconception that the tambalacoque absolutely required the dodo for its seeds to germinate. Plant Science Bulletin. 50: 105-108.

Het Tweede Boeck. (1601). Journal oft Dagh-register / inhoundende een warachtig verhael ende historische vertellinghe van de Reyse/ gedaen door de acht shepen van Amstelredame / gheseylt in den Maert Martij 1598 onder 't beleydt van den Admirael Jacob Cornelisz. Neck ende Wybrandt van Warwijck als vice admirael –van hare zeylagie ende gedenwaerdighe zaken ende geschiedenissen haer op de voortz-reys bejeghent. Ghedruct tot Amstelredam by Cornelis Claesz. Opt Water in 't Schrijf-boeck. Amsterdam.

van Heteren, Anneke H., van Dierendonk, R. C. H., van Egmond, M. A. N. E., ten Hagen, S. L. and Kreuning, J. (2017). Neither slim nor fat: estimating the mass of the dodo ([i]Raphus cucullatus[/i], Aves, Columbiformes) based on the largest sample of dodo bones to date. PeerJ 5:e4110.

Heupink, Tim H., van Grouw, Hein and Lambert, David M. (2014). The mysterious Spotted Green Pigeon and its relation to the Dodo and its kindred. BMC Evolutionary Biology 14: 136.

Hickey, Georgina. (1988). Alice, the Dodo and the Tambalacoque Tree. Australian Natural History 22(8): 356-357.

Horn, B. K. P. 1978. Dodo apocrypha. Science News 113: 19.

Horn, Ivo R. et al. (2019). Palaeoproteomics of bird bones for taxonomic classification. Zoological Journal of the Linnean Society. [Abstract]

del Hoyo, J., Collar, N.J., Christie, D.A., Elliott, A. and Fishpool, L.D.C. 2014. HBW and BirdLife International Illustrated Checklist of the Birds of the World. Volume 1: Non-passerines. Lynx Edicions BirdLife International, Barcelona, Spain and Cambridge, UK.

Hume, Julian Pender. (2003). The journal of the flagship Gelderland – dodo and other birds on Mauritius 1601. Archives of Natural History 30: 13-27.

Hume, Julian Pender. (2005). Contrasting taphofacies in ocean island settings: the fossi! record of Mascarene vertebrates. In ALCOVER, J .A. & BOVER, P. (eds.): Proceedings of the International Symposium "Insular Vertebrate Evolution: the Palaeontological Approach': Monografies de la Societat d'Història Natural de les Balears 12: 129-144.

Hume, Julian Pender. (2006). The history of the Dodo [i]Raphus cucullatus[/i] and the penguin of Mauritius. Historical Biology 18(2): 65-89.

Hume, Julian Pender. (2009a). The great Dodo dispute. BBC History 10 (3): 50-53.

Hume, Julian Pender. (2009b). In search of the Dodo. Darwin 200 BBC Knowledge 4: 26-31.

Hume, Julian Pender. (2012). The Dodo: from extinction to the fossil record. Geology Today 28(4): 147-151. [Abstract]

Hume, Julian Pender. (2013). A synopsis of the pre-human avifauna of the Mascarene Islands, pp. 195-237. In: Göhlich UB, Kroh A, editors. Proceed. 8th Internat. Meeting Society ofAvian Paleontology and Evolution. Pp.195–237. Wien,  Naturhistorisches Museum.

Hume, Julian Pender and Cheke, Anthony S. (2004). The White Dodo of Réunion Island: unravelling a scientific and historical myth. Archives of Natural History 31(1): 57-79.

Hume, Julian Pender, Cheke, Anthony S. and McOran-Campbell, A. (2009). How Owen ‘stole’ the Dodo: academic rivalry and disputed rights to a newly-discovered subfossil deposit in nineteenth century Mauritius. Historical Biology 21(1-2): 33-49. [Abstract]

Hume, Julian Pender, Datta, A. and Martill, D. M. (2006). Unpublished drawings of the dodo Raphus cucullatus and notes on dodo skin relics. Bull B.O.C. 126A: 49-54.

Hume, Julian Pender et al. (2014). Rediscovery of a lost Lagerstätte: a comparative analysis of the historical and recent Mare aux Songes dodo excavations on Mauritius. Historical Biology: An International Journal of Paleobiology DOI: 10.1080/08912963.2014.945927 [Abstract]

Hume, Julian Pender, Martill, D. M. and Dewdney, C. (2004). Dutch diaries and the demise of the dodo. Nature 429: 621.

Hume, Julian Pender et al. (2014). In the footsteps of the bone collectors: nineteenth-century cave exploration on Rodrigues Island, Indian Ocean. Historical Biology DOI:10.1080/08912963.2014.886203 [Abstract]

Hume, Julian Pender and Taylor, Christine. (2016). A gift from Mauritius: William Curtis, George Clark and the Dodo. J Hist Collections. doi: 10.1093/jhc/fhw033 [Abstract]

Hume, Julian Pender and Winters, Ria. (2015). Captive birds on Dutch Mauritius: bad-tempered parrots, warty pigeons and notes on other native animals. Historical Biology. DOI:10.1080/08912963.2015.1036750 [Abstract]

Iredale, Tom. (1924). As extinct as the Dodo. The Australian Museum Magazine 2(4): 117-120.

Iverson, J. B. (1987). Tortoises, not dodos, and the tambalacoque tree. J. Herpetol. 21: 229-230.

Iwanow, A. (1958). An Indian picture of the Dodo. Journal of Ornithology 99(4): 438-440.

Jackson, Andrew. (2013). Added credence for a late Dodo extinction date. Historical Biology 26(6): 699-701. [Abstract]

Jango, A. (2005), ‘Discovery of isolated dodo bones (Raphus cucullatus(L.) Aes, Columbiformes) from Mauritius cave shelters highlights human predation, with a comment on the status of the family Raphidae Wtmore, 1930.’ Annales de Paleontologie 91: 167-180. [Abstract]

Janoo, Anwar. (1996). On a Hitherto Undescribed Dodo Cranium, Raphus cucullatus L. (Aves, Columbiformes), with a brief Taxonomical Overview of this Extinct Flightless Mascarene Island Bird. Bulletin du Museum National d’Histoire Naturelle Section C Sciences de la Terre Paleontologie Geologie Mineralogie 18: 57-77.

Janoo, Anwar. (2000). Rooting the Dodo Raphus cucullatus Linnaeus 1758 and the Solitaire Pezophaps solitaria Gmelin 1789 within the Omithurae: a cladistic reappraisal. Ostrich 71(1-2): 323-329. [Abstract]

Janoo, Anwar. (2005). Discovery of isolated dodo bones [ Raphus cucullatus (L.), Aves, Columbiformes] from Mauritius cave shelters highlights human predation, with a comment on the status of the family Raphidae Wetmore, 1930. Annales De Paleontologie 91(2): 167-180. [Abstract]

Johnson, K. P. and Clayton, D. H. (2000). Nuclear and mitochondrial genes contain similar phylogenetic signals for pigeons and doves (Aves: Columbiformes). Molecular Phylogenetics and Evolution 14(1): 141-151.

Jouanin, Christian. (1962). Inventaire des oiseaux éteints ou en voie d’extinction conservés au Muséum de Paris. Terre et Vie 109: 275-301.

Kalla, Abdool Cader. (unpublished?). 150 years of Dodo bones: deceit, greed and revelation.

Kallio, Harri. (2004). The Dodo and Mauritius Island: imaginary encounters. London: Dewi Lewis Publishing.

King, H.C. (1946). Interim Report on Indigenous Species in Mauritius. Port Luis, Mauritius: Government Printer.

Kitchener, Andrew C. (1993a). On the external appearance of the dodo, Raphus cucullatus (L., 1758). Archives of Natural History 20(2): 279-301.

Kitchener, Andrew C. (1993b). Justice at last for the dodo. New Scientist (28 August).

Knox, Alan G. and Walters, Michael P. (1994). Extinct and endangered birds in the collections of The Natural History Museum. British Ornithologists' Club Occasional Publications 1: 1-292. [p. 134]

Lawrence, Natalie. (2015). Assembling the dodo in early modern natural history. The British Journal for the History of Science 48(3): 387-408. [Abstract]

Lawton, Graham. (2019). The last dodo. New Scientist 244(3261): 74-76. [Abstract]

Ley, Willy. (1959). Exotic Zoology. New York: Viking. [pp. 334-354]

Linnaeus, Carolus. (1766). Systema Naturae per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Edition 12, volume 1. Stockholm: Laurentii Salvii.

Livezey, Bradley C. (1993). An ecomorphological review of the dodo (Raphus cucullatus) and solitaire (Pezophaps solitaria), flightless Columbiformes of the Mascarene Islands. Journal of Zoology 230(2): 247-292. [Abstract]

Louchart, Antoine and Mourer-Chauviré, Cécile. (2011). The dodo was not so slim: leg dimensions and scaling to body mass. Naturwissenschaften 98(4): 357-358.

Lüttschwager, J. (1961). Die Drontevögel. Die Neue Brehm-Bücherei 1961: 276.

McMichael, Donald F. (1967). The conservation of animals and plants. Australian Natural History 15(12): 423-428.

McNaughton, S. J. 1989. Ecosystems and conservation in the twenty-first century. - In: Western, D. and Pearl, M. (eds), Conservation for the twenty-first century. Oxford Univ. Press, Oxford, pp. 109-120.

Meijer, H. J., Gill, A., Louw, P. G., Hoek Ostende, L. W., Hume, J. P. and Rijsdijk, K. F. (2012). Dodo remains from an in situ context from Mare aux Songes, Mauritius. Naturwissenschaften -Dusseldorf- 99(3): 177-184. [Abstract]

Middleton Gregory J. and Hume, Julian Pender. (2016). The discovery of a Dodo [i]Raphus cucullatus[/i] Linn. (Aves, Columbiformes) in a highland Mauritian lava cave. Helictite 42: 13-20.

Mlíkovský, J. (2004). Extinction of the Dodo Raphus cucullatus (Aves: Raphidae): Dating Reconsidered. Journal of the National Museum, Natural History Series 173: 111-112.

Mlíkovský J., 2010a: Dodo císaře Rudolfa II. – The dodo of Emperor Rudolf II. – In: Kotková O. (ed.): Roelandt Savery: malíř ve službách císaře Rudolfa II. – Roelandt Savery: a painter in the services of Emperor Rudolf II. Pp. 325-331. Praha: Národní galerie v Praze. [Parallel text in Czech and English.]

Mlíkovský, Jiří. (2010b). De dodo van keizer Rudolf II. – In: De Jaegere I. & Kotková O. (eds.): Roelandt Savery 1576-1639. Pp. 51-53. Kortrijk: Snoeck. [Dutch translation of Mlíkovský 2010a.]

Mlíkovský, Jiří. (2012). Extinct and nearly extinct birds in the collections of the National Museum, Prague, Czech Republic. Journal of the National Museum (Prague), Natural History Series 181(9): 95-123. [automatic download]

Mlíkovský J., Benda P., Moravec J. & Šanda R., 2011: Type specimens of recent vertebrates in the collections of the National Museum, Prague, Czech Republic. Journal of the National Museum (Prague), Natural History Series 180: 137-168.

Moree, P. (1998). A concise history of Dutch Mauritius, 1598-1710. London and New York: Kegan Paul International.

Myers, N. (ed.). 1985. The Gaia atlas of planet management. Pan Books, London.

Neville, H. (1668). The Isle of Pines, or, a late discovery of a fourth Island, in Terra Australis Incognita. London: Printed for Allen Banks and Charles Harper.

Newton, A. 1904. Further discovery of dodos’ bones. Nature 70:626.

Newton, Alfred. (2009). On a Living Dodo shipped for England in the Year 1628. Journal of Zoology 42(1): 447-449. [published in 1874?]

Newton, E. (1890). On the reported discovery of Dodo's bones in a cavern in Mauritius. Proc. Zool. Soc. Lond. 1890: 402-403.

Newton, E. and Gadow, H. (1893). IX. On additional bones of the dodo and other extinct birds of Mauritius obtained by Mr. Theodore Sauzier. The Transactions of The Zoological Society of London 13(7): 281-302.

Nicholls, Henry. (2006). Ornithology: Digging for dodo. 443(7108): 138-140. [Abstract]

Nieremberg, J. E. (1635). J. E. Nieremberg...Historia Naturae maxime peregrinae, libris XVI, distincta In quibus rarissma Naturae arcana & ignota Indiarum Animalia...Antverpiae. Antwerp.

Nowak-Kemp, M. and Hume, Julian Pender. (2016a). The Oxford Dodo. Part 1: the museum history of the Tradescant Dodo: ownership, displays and audience. Historical Biology. DOI: 10.1080/08912963.2016.1152471

Nowak-Kemp, M. and Hume, Julian Pender. (2016b). The Oxford Dodo. Part 2: from curiosity to icon and its role in displays, education and research. Historical Biology. DOI: 10.1080/08912963.2016.1155211

Olson, S. L. (1989). Extinction on islands: man as a catastrophe, pp. 50-53. In: Western, D. and Pearl, M. C. (eds.). Wildlife Conservation International. New York Zoological Society, Oxford University Press, NY.

Oudemans, A. C. (1917). Dodo-studien, naar aanleidingvan de vondst van een gerelsteen met Dodo-beeld van 1561 to Vere. 19. Amsterdam: Verhandelingen der Koninklijke Akademic van Wetenschappen. pp. 1-140.

Ovenell, R. F. (1992). The tradescant dodo. [i]Arch. Nat. Hist.[/i] [b]19[/b]: 145-152.

Owadally, A. W. (1979). The Dodo and the tambalacoque tree. Science 203(4387): 1363-1364.

Owen, Richard. (1866). Memoir on the dodo (Didus ineptus), with an historical introduction by the late W.J. Broderip. London (UK): Taylor & Francis. vi, 56 pp.

Owen, Richard. (1872). On the Dodo (Part II). Notes on the Articulated Skeleton of the Dodo (Didus ineptus, Linn.) in the British Museum. 7: 513-525. [513-526 plates 64-66?]

Parish, Jolyon C. (2012). Bibliography of the Didinæ, in The Dodologist’s Miscellany. Available at: [accessed 9 February 2017] 

Parish, Jolyon C. (2013). The Dodo and the Solitaire: a natural history. Indianapolis: Indiana University Press, 406 pp.

Parish, Jolyon C. and Cheke, Anthony S. (2018). A newly-discovered early depiction of the Dodo (Aves: Columbidae: Raphus cucullatus) by Roelandt Savery, with a note on another previously unnoticed Savery Dodo. Historical Biology. [Abstract]

Pickering, J. (2010). The Oxford Dodo: The sad story of the ungainly bird that became an Oxford icon. Oxford University Museum of Natural History. 32 pp.

Pimm, Stuart. (2003). Expiry dates. Nature 426(6964): 235.

Pinto-Correia, Clara. (2003). Return of the crazy bird: the sad, strange tale of the Dodo. New York: Copernicus Books.

Pitot, A. 1905. T’Eylandt Mauritius. Esquisses historiques (1598–1710). Pre´ce´de´s d’une notice sur la de´couverte des Mascareignes et suivies d’une monographie du Dodo, des Solitaires de Rodrigue et de Bourbon et de l’Oiseau Bleu. Port Louis (Mauritius): Coignet Fre`res & Cie. xv þ 372 p.

Randall A, Pepe A, Yamartino K, Kimelblatt A, Claeesens L. 2014. How fat was the Dodo? The first mass estimate from digital body reconstruction based on a complete skeleton of Raphus cucullatus. In: Society of vertebrate paleontology annual meeting 2014. Berlin Poster Abstract.

Renshaw, G. (1931). The Mauritius dodo. Journ. Soc. Preservation Fauna Empire (n.s.), part 15: 14-21.

Renshaw, G. (1933). The dodo and aphanapteryx. Nature 131: 728.

Reuss, A. E. (1855). Paläontologische Miscellen. I. Über ein Schädelfragment der Dronte im Prager Museum. – Denkschriften der Kaiserlichen Akademie der Wissenschaften, MathematischNaturwissenschaftliche Classe 10: 71-78.

Rheinhardt, J. T. (1842). Nojere Oplysning om det i Kjoben-havn funde Dronte hoved. Naturhistorisk Tidskrift 4: 71-72.

Ricci, Vincenzo and Özçakar, Levent. (2018). The Dodo Bird is not Extinct: Ultrasound Imaging for Supraspinatus Tendinosis. American Journal of Physical Medicine & Rehabilitation. doi: 10.1097/PHM.0000000000000992

Richon, Emmanuel and Winters, Ria. (2014). The intercultural dodo: a drawing from the School of Bundi, Rājasthān. Historical Biology.

Rijsdijk, K. F., Hume, J. P., Bunnik, F., Florens, F. B., Baider, C., Shapiro, B., van der Plicht, J., Janoo, A., Griffiths, O. and van den Hoek Ostende, L. W. (2009). Mid-Holocene vertebrate bone Concentration-Lagerstatte on oceanic island Mauritius provides a window into the ecosystem of the dodo (Raphus cucullatus). Quaternary Science Reviews  28(1-2): 14-24. [Abstract]

Rijsdijk, K. F., J. P. Hume, P. G. B. de Louw, H. J. M. Meijer, A. Janoo, E. J. de Boer, L. Steel, J. de Vos, L. G. van der Sluis, H. Hooghiemstra, F. B. V. Florens, C. Baider, T. J. J. Vernimmen, P. Baas, A. H. van Heteren, V. Rupear, G. Beebeejaun, A. Grihault, J. van der Plicht, M. Besselink, J. K. Lubeek, M. Jansen, S. J. Kluiving, H. Hollund, B. Shapiro, M. Collins, M. Buckley, R. M. Jayasena, N. Porch, R. Floore, F. Bunnik, A. Biedlingmaier, J. Leavitt, G. Monfette, A. Kimelblatt, A. Randall, P. Floore, and L. P. A. M. Claessens. (2015). A review of the dodo and its ecosystem: insights from a vertebrate concentration Lagerstätte in Mauritius, pp. 3–20 in L. P. A. M. Claessens, H. J. M. Meijer, J. P. Hume, and K. F. Rijsdijk (eds.), Anatomy of the Dodo (Raphus cucullatus L., 1758): An Osteological Study of the Thirioux Specimens. Society of Vertebrate Paleontology Memoir 15. Journal of Vertebrate Paleontology 35(6): supplement 1, 3-20.

Rijsdijk, K. F., J. Zinke, P. G. B. de Louw, J. P. Hume, J. van der Plicht, H.Hooghiemstra, H. J. M. Meijer, H. B. Vonhof, N. Porch, F. B. V. Flo-rens, C. Baider, B. van Geel, J. Brinkkemper, T. Vernimmen, and A.Janoo. 2011. Mid-Holocene (4.2 kyr BP) mass mortalities in Mauri-tius (Mascarenes): insular vertebrates resilient to climatic extremesbut vulnerable to human impact. The Holocene 21:1179–1194.

Roberts, D. L. (2013). Refuge-Effect Hypothesis and the Demise of the Dodo. Conservation Biology 27: 1478-1480. [Abstract]

Roberts, David L. and Solow, Andrew R. (2003). Flightless birds: When did the dodo become extinct? Nature 426(6964): 245. [Abstract]

Rothschild, Lionel Walter. (1907). Extinct birds: an attempt to write in one volume a short account of those birds which have become extinct in historical times, that is within the last six or seven hundred years: to which are added a few which still exist, but are on the verge of extinction. London: Hutchinson & Co. XXIX + 243 pp. [p. 172-174, pl. 24-24C]

Sayol, Ferran, Steinbauer, Manuel J., Blackburn, Tim M., Antonelli, Alexandre and Faurby, Søren. (2020). Anthropogenic extinctions conceal widespread evolution of flightlessness in birds. Science Advances 6(49): eabb6095. [Supplementary Material (Data File S1)]

Schlegel, H. (1854). Ook een Woordje over den Dodo (Didus ineptus) en zijne Verwanten. Verslagen en Mededeelingen der Koninklijke Akademie van Wetenschappen 2: 232-256.

Serrano, Francisco J. et al. (2020). Morphological Disparity of the Humerus in Modern Birds. Diversity 12(5): 173.

Servaas, A. J. (1887). Hongersnood in Suratta a° 1631, enz. Waarheid of leugen mag men wel vragen aangaande 'tgeen volgt. De Navorscher 37(1): 4-8.

Shapiro, Beth, Sibthorpe D, Rambaut A, Austin J, Wragg GM, Bininda-Emonds ORP, Lee PLM, Cooper A. (2002). Flight of the Dodo. Science 295(5560): 1683.

Shuker, Karl P. N. (1989). How Dead Is the Dodo? Fate 42: 62-69.

Shuker, Karl P. N. (1998). From Dodos to Dimetrodons. Strange Magazine 19: 22.

Shuker, Karl P. N. (1999). Mysteries of Planet Earth: An Encyclopedia of the Inexplicable. Carlton Books. 192 pp.

Shuker, Karl P. N. (2012, 22 September). The stuffed dodo that was a no-no! Blog post at shukernature, available at:

Soares, André E. R. et al. (2016). Complete mitochondrial genomes of living and extinct pigeons revise the timing of the columbiform radiation. BMC Evolutionary Biology 16: 230.

Stiles, E. W. 1989. Fruits, seeds, and dispersal agents. In: Abrahamson, W. G. (ed.), Plant-animal interactions. McGraw-Hill, NY, pp. 87-113.

Stresemann, Erwin. (1958). Wie hat die Dronte (Raphus cucullatus L.) ausgesehen? Journal of Ornithology 99(4): 441-459.

Storer, R. W. (1970). Independent evolution of the Dodo and the Solitaire. Auk 87: 396-370.

Strickland, Hugh Edwin. (1844). On the evidence of the former existence of struthious birds distinct from the Dodo in the islands near Mauritius. Proc Zool Soc Lond. 1844/12: 77-79.

Strickland, Hugh Edwin. (1850). Supplementary notices regarding the Dodo and its kindred. 9. Discovery of a third example of a Dodo’s head. Annals and Magazine of Natural History (2) 6: 290.

Strickland, Hugh Edwin and Melville, Alexander Gordon. (1848). The dodo and its kindred: Or the history, affinities, and osteology of the dodo, solitaire, and other extinct birds of the islands Mauritius, Rodriguez and Bourbon. London (UK): Reeve, Benham & Reeve. viii, 141 pp. [WARNING: 14 MB file]

Teixeira, Dante Martins. (2019). The German painter Carl Borromäus Andreas Ruthart (ca. 1630-1703) and some still unregistered images of the extinct dodo, Raphus cucullatus (Linnaeus, 1758) (Aves, Columbiformes). Arquivos De Zoologia 50(4): 191-200.

Temple, R. C. (ed.). (1914). The travels of Peter Mundy in Europe and Asia, 1608-1667. Hakluyt Society: SII, 35(2): 1-437. London.

Temple, Stanley A. (1977). Plant-Animal Mutualism: Coevolution with Dodo Leads to Near Extinction of Plant. Science, New Series, 197(4306): 885-886.

Temple, S.A. (1979). The dodo and the tambalacoque tree. Science, 203(4387), 1364.

Tradescant, J. (1656). Museum Tradescantianum, or a collection of rarities preserved at South Lambeth, near London.

Turvey, Samuel T. and Cheke, Anthony S. (2008). Dead as a dodo: the fortuitous rise to fame of an extinction icon. Historical Biology 20(2): 149-163.

Tyrberg, Tommy. (2009). Holocene avian extinctions, pp. 63-106. In: Turvey, Samuel T. (ed.). Holocene Extinctions. Oxford, UK & New York, USA: Oxford University Press. xii + 352 pp.

Valledor de Lozoya A., 2003: An unnoticed painting of a white dodo. – Journal of the History of Collections 15: 201-210.

Vargas, Pablo. (2023). Exploring ‘endangered living fossils’ (ELFs) among monotypic genera of plants and animals of the world. Frontiers in Ecology and Evolution 11: 1100503.

Vaughan, R. E. (1984). [untitled letter]. - Animal Kingdom 87: 6-7.

Vaughn, R.E. and Wiehe, P.O. (1937). Studies on the vegetation of Mauritius I. A preliminary survey of the plant communities. Journal of Ecology, 25, 289-243.

Vaughn, R.E. and Wiehe, P.O. (1941). Studies on the vegetation of Mauritius III. The structure and development of the upland forest. Journal of Ecology, 29, 127-160.

Warnett, J. M. et al. (2021). The Oxford Dodo. Seeing more than ever before: X-ray micro-CT scanning, specimen acquisition and provenance. Historical Biology 33(10): 2247-2255.

Webster, L. M. I. and McEwing, R. (In Press, 2013). Resurrecting the Dodo: Positive control DNA for species identification Forensic Science International: Genetics Supplement Series. [Abstract]

Winters, Ria and Hume, Julian Pender. (2014). The dodo, the deer and a 1647 voyage to Japan. Historical Biology. DOI:10.1080/08912963.2014.884566

Winters, Ria and Hume, Julian Pender and Leenstra, M. (2017). A famine in Surat in 1631 and Dodos on Mauritius: a long lost manuscript rediscovered. Archives of Natural History 44(1): 134-150. [Abstract]

Wissen, B. van (ed.). (1995). Dodo Raphus cucullatus. Amsterdam (The Netherlands): Zoölogisch Museum, Amsterdam University. 102 p.

Witmer, Mark C. and Cheke, Anthony S. (1991). The dodo and the tambalacoque tree: an obligate mutualism reconsidered. Oikos 61(1): 133-137.

Wood, Casey A. (1927). Two hitherto Unpublished Pictures of the Mauritius Dodo. Ibis 69(3): 724-732.

Zinoview, Andrei V. (2014). Preliminary observations on hindlimb myology and syndesmology of the Dodo (Raphus cucullatus, Columbiformes). / eds. Delfino M., Carnevale G., Pavia M. Abstract book and field guide, XII Annual Meeting of EAVP. Museo Regionale di Scienze Naturali. Torino: EAVP. P. 159.

Ziswiler, V. (1996). Der Dodo. Fantasien und fakten zu einem vershwunderen Vogel. Zu¨rich (Switzerland): Zoologisches Museum der Universta¨t Zu¨rich. 94 p.

Extinct, tv show, season 1, episode 1, "Dodo", originally aired 25 September 2001, see here:

Wild Kratts, season 3, episode 25, Back in Creature Time, Part 1: Day of the Dodo, 7 July, 2014, full episode available here:


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