Gopherus hexagonatus Cope, 1893



Taxonomy & Nomenclature

Synonyms: Testudo campester Hay, 1908; Bysmachelys canyonensis Johnston, 1937; Gopherus canyonensis Johnston, 1937; Gopherus huecoensis Strain, 1966


Conservation Status


Last record: Terminal Pleistocene (Esker et al., 2019)



Texas, USA


Biology & Ecology










Original scientific description:

Cope, Edward Drinker. (1893). A preliminary report on the vertebrate paleontology of the Llano Estacado. Fourth Annual Report of the Geological Survey of Texas 1892: 11-136.


Other references:

Esker, Donald A. (2019). Thermoregulation and Dental Isotopes Reveal the Behavior and Environment of Pleistocene Megafauna at Waco Mammoth National Monument. PhD thesis, Baylor University.

Esker, Donald A., Forman, Steve L. and Butler, Dava K. (2019). Reconstructing the mass and thermal ecology of North American Pleistocene tortoises. Paleobiology 45(2): 363-377. [Abstract]


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