Tokudaia muenninki (Johnson, 1946)

Okinawa spiny rat, Muennink's spiny rat



Taxonomy & Nomenclature



Conservation Status

Last record: 1978

Rediscovered in 2008

IUCN status: Critically Endangered



Yanbaru area (i.e. northern part), Okinawa Island, Ryukyu Islands, Japan


Biology & Ecology










Original scientific description:

Johnson, D. H. (1946). The spiny rat of the Riu Kiu island. Proceedings of the Biological Society of Washington 59: 169-172.


Other references:

Abe, H., Ishii, N., Ito, T., Kaneko, Y., Maeda, K., Miura, S. and Yoneda, M. 2005. A Guide to the Mammals of Japan. Tokai University Press, Kanagawa, Japan.

Fisher, Diana O. (2011). Trajectories from extinction: where are missing mammals rediscovered? Global Ecology & Biogeography 20: 415-425. [Appendix S1]

Ishii, N. and Kaneko, Y. (2008). Tokudaia muenninki. The IUCN Red List of Threatened Species. Version 2014.3. ( Downloaded on 07 May 2015.

Kaneko, Y. 2001. Morphological discrimination of the Ryukyu spiny rat (genus Tokudaia) between the islands of Okinawa and Amami Oshima, in the Ryukyu Islands, southern Japan. Mammal Study 26(1): 17-33.

Ministry of the Environment. 2007. The revised Japanese Red List.

Scheffers, Brett R., Yong, Ding Li, Harris, J. Berton C., Giam, Xingli and Sodhi, Navjot S. (2011). The world’s rediscovered species: back from the brink? PLoS ONE 6(7): e22531. [Supporting Information (Table S1)]

Yamada, Fumio et al. (2010). Rediscovery After Thirty Years Since the Last Capture of the Critically Endangered Okinawa Spiny Rat Tokudaia muenninki in the Northern Part of Okinawa Island. Mammal Study 35(4): 243-255. [Abstract]


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