Eupetaurus cinereus Thomas, 1888:258

Woolly flying squirrel, Wooly flying squirrel, Giant woolly gliding squirrel



Taxonomy & Nomenclature



Conservation Status

Last Record: 1924

Rediscovered in 1994

IUCN RedList status: Endangered



Kashmir, Pakistan


Biology & Ecology










Original scientific description:

Thomas, O. (1888). Eupetaurus-A new form of flying squirrel from Kashmir. J. Asiatic Soc. Bengal 57: 256-260.


Other references:

Agrawal, V. C. (2000). Taxonomic studies on Indian Muridae and Hystricidae (Mammalia: Rodentia). Records of the Zoological survey of India 180: 1-177.

Agrawal, V.C,. and Chakraborty, S. 1970. Occurrence of the Woolly Flying Squirrel (Eupetaurus cinereus Thomas (Mammalia: Rodentia: Sciuridae) in North Sikkim. Journal of the Bomaby Natural History 66: 615-616.

Agrawal, V. C. and Chakraborty, S. 1979. Catalogue of mammals in the Zoological Survey of India. Part 1. Sciuridae. Records of the Zoological Survey of India 74: 333-481.

Corbet, G. B. and Hill, J. E. (1992). Mammals of the Indo-Malayan Region: a Systematic Review. Oxford University Press, Oxford, UK.

Dar, N. M. (1996). Survey of woolly flying squirrel in northern Pakistan. Final report submitted to WWF-Pakistan. 14 pp.

Din, Jaffar Ud et al. (2015). Note on the Giant Woolly Gliding Squirrel Eupetaurus cinereus (Mammalia: Rodentia: Sciuridae) in northern Pakistan. Journal of Threatened Taxa 7(9): 7602-7604.

Fisher, Diana O. (2011). Trajectories from extinction: where are missing mammals rediscovered? Global Ecology & Biogeography 20: 415-425. [Appendix S1]

Hasan, L. 2008. An anatomy of state failures in the forest management in Pakistan. MPRA Paper No. 6513. Munich Personal RePEc Archive.

Hoffman, R. S., Anderson, C. G., Thorington Jr., R. W. and Heaney, L. R. (1993). Family Sciuridae, pp. 419-465. In: Wilson, D. and Reeder, D. M. Mammal Species of the World, 2nd edition. Washington: Smithsonian Institute Press. 1206 pp.

Jackson, Stephen M. et al. (2021). Across the great divide: revision of the genus Eupetaurus (Sciuridae: Pteromyini), the woolly flying squirrels of the Himalayan region, with the description of two new species. Zoological Journal of the Linnean Society.

McKenna, M. C. (1962). Eupetaurus and the living petauristine sciurids. American Museum Novitates 2104: 38 pp.

Mirza, Z. B. (2005). The final report of the post cranila skeltetal measurements of Wooly Flying Squirell Eupetaurus cinereus with observations of its behaviour. Proceedings of International Zoolgical Congress, 2004. Zoolgical society of Pkaistan, Punjab University Lahore.

Molur, S., Srinivasulu, C., Srinivasulu, B., Walker, S., Nameer, P. O. and Ravikumar, L. 2005. Status of non-volant small mammals: Conservation Assessment and Management Plan (C.A.M.P) workshop report. Zoo Outreach Organisation / CBSG-South Asia., Comibatore, India.

Nowak, R.M. 1999. Walker's Mammals of the World. The Johns Hopkins University Press, Baltimore, USA and London, UK.

OSHIDA, T., SHAFIQUE, C.M. AND BARKAKTI, S. (2005). A note on a new habitat of the woolly flying squirrel Eupetaurus cinereus in the northern area of Pakistan. Mammal Study 30: 73-76.

Pal, Ranjana et al. (2018). Range extension and high elevation record for the endangered woolly flying squirrel Eupetaurus cinereus in Western Himalaya, India. Mammalia. doi: [Abstract]

Pal, Ranjana, Bhattacharya, Tapajit and Sathyakumar, S. (2020). Woolly Flying Squirrel Eupetaurus cinereus: a new addition to the diet of Snow Leopard Panthera uncia. J. Bombay Nat. Hist. Soc. 117. doi: 10.17087/jbnhs/2020/v117/142056

Pradhan, M. S. and Talmale, S. S. (2009). List of valid Rodent taxa (Class: Mammalia, Order: Rodentia) from Indian Subcontinent region including Myanmar. Published by the Director, Zool. Surv. India, Kolkata. Rec. zool. Surv. India, Kolkata. 297: 1-239.

Qamar, Qamar Zaman et al. (2012). New Distribution Information on Woolly Flying Squirrel (Eupetaurus cinereus Thomas, 1888) in Neelum Valley of Azad Jammu and Kashmir, Pakistan. Pakistan J. Zool. 44(5): 1333-1342.

Roberts, T. J. (1977). The Mammals of Pakistan. London: Ernest Benn.

Roberts, T. J. (1997). The mammals of Pakistan. Revised edition. Oxford University Press, Karachi. 525 pp.

Schaub, S. (1953). Remarks on the distribution and classification of the "Histricomorpha". Verhandlunden naturforchenden Gesellschaft Basel 64: 389-400.

Scheffers, Brett R., Yong, Ding Li, Harris, J. Berton C., Giam, Xingli and Sodhi, Navjot S. (2011). The world’s rediscovered species: back from the brink? PLoS ONE 6(7): e22531. [Supporting Information (Table S1)]

SHAFIQUE, C. M., BARKATI, S., OSHIDA, T. AND ANDO, M. (2006). Comparison of diets between two sympatric flying squirrel species in Northern Pakistan. J. Mammal. 87: 784-789.

Sheikh, K. M. and Molur, S. (eds.). (2004). Status and Red List of Pakistan’s Mammals. Based on the Conservation Assessment and Management Plan. 312pp. IUCN Pakistan.

SHEIKH, K. M. AND MOLUR, S. (eds.). (2005). Status and Red list of Pakistan’s Mammals. Based on conservation assessment and management planning for mammals. IUCN- Pakistan. 344 pp.

Srinivasulu, C., Chakraborty, S. and Pradhan, M. S. (2004). Checklist of sciurids (Mammalia: Rodentia: Sciuridae) of South Asia. Zoos’ Print Journal 19(2): 1351-1360.

Walters, M. J. (1995). Adventures of a Flying Squirrel. Animals, July/Aug.

YU, F., 2002. Systematics and biogeography of flying squirrels in the Eastern and the Western Trans-Himalayas. Unpublished Ph.D. dissertation, University of Florida. USA. 228 pp.

Zahler, P. (1996). Rediscovery of the woolly flying squirrel (Eupetaurus cinereus). Journal of Mammalogy 77(1): 54-57.

Zahler, P. (2001). The woolly flying squirrel and gliding: Does size matter? Acta Theriol. 46: 429-435.

Zahler, P. (2010). Eupetaurus cinereus. The IUCN Red List of Threatened Species. Version 2015.2. ( Downloaded on 05 July 2015.

Zahler, P. and Khan, M. 2003. Evidence for dietary specialization on pine needles by the woolly flying squirrel (Eupetaurus cinereus). Journal of Mammalogy 84(2): 480-486.

Zahler, P. and Woods, C.A. 1997. The status of the woolly flying squirrel (Eupetaurus cinereus) in Northern Pakistan. In: S.A. Mufti, C.A. Woods and S.A. Hasan (eds), Biodiversity of Pakistan, pp. 485-514. Florida Museum of Natural History, Gainesville, Florida.


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