Huperzia nutans (Brack.) Rothm. (1944:62)

Nodding clubmoss, Wawaeʻiole

Taxonomy & Nomenclature

Synonyms: Lycopodium nutans Brack. in U.S. Expl. Exped., Filic. 16: 327 (1854); Phlegmariurus nutans (Brack.) W.H.Wagner in Contr. Univ. Michigan Herb. 20: 241 (1995)

Conservation Status

Rediscovered [by implication]

Listed as Extinct by (WCMC, 1992:215), but known to be extant today (NatureServe, 2025).

Distribution & Habitat

Hawaiian Islands, USA (& Vanuatu?)

Anatomy & Morphology

Biology & Ecology

Hypodigm

Media

References

Bruegmann, M. M. and Caraway, V. (2003). Phlegmariurus nutans. The IUCN Red List of Threatened Species 2003: e.T44062A10847463. https://dx.doi.org/10.2305/IUCN.UK.2003.RLTS.T44062A10847463.en. Accessed on 14 August 2025.

Lorence, D. H., Keir, M., Nyberg, B. and Wood, K. (2023). Phlegmariurus nutans. The IUCN Red List of Threatened Species 2023: e.T44062A83804757. https://dx.doi.org/10.2305/IUCN.UK.2023-1.RLTS.T44062A83804757.en. Accessed on 14 August 2025.

NatureServe. (2025). Huperzia nutans: Nodding Clubmoss. NatureServe Network Biodiversity Location Data accessed through NatureServe Explorer [web application]. NatureServe, Arlington, Virginia. Available at: https://explorer.natureserve.org/ [Accessed 14 August 2025]

Palmer, D. D. (2003). Hawai'i's Ferns and Fern Allies. University of Hawai'i Press, Honolulu.

Plunkett, G. M., Ranker, T. A., Sam, C. and Balick, M. J. (2022). Towards a checklist of the vascular flora of Vanuatu. Candollea 77: 105-118.

POWO. (2025). Plants of the World Online (online resource). Facilitated by the Royal Botanic Gardens, Kew, United Kingdom. Available at: https://powo.science.kew.org/ [Accessed 14 August 2025]

Ranker, T. A., Imada, C. T., Lynch, K., Palmer, D. D., Vernon, A. L. and Thomas, M. K. (2019). Taxonomic and nomenclatural updates to the fern and lycophyte flora of the Hawaiian Islands. American Fern Journal 109(1): 54-72.

Roskov Y. et al. (eds.). (2018). World Ferns: Checklist of Ferns and Lycophytes of the World. Species 2000 & ITIS Catalogue of Life Naturalis, Leiden, the Netherlands.

U.S. Fish and Wildlife Service, Pacific Region. (1997). Draft recovery plan for Oahu plants. U.S. Fish and Wildlife Service, Portland, Oregon.

U.S. Fish and Wildlife Service. (2019). Endangered and threatened wildlife and plants; initiation of 5-year status reviews for 91 species in Oregon, Washington, Hawaiʻi, and American Samoa.

Wagner, W., Herbst, D. and Sohmer S. (1999). Manual of the Flowering Plants of Hawaii. Bernice Pauahi Bishop Museum Special Publication 91: 1-1918.

WCMC (World Conservation Monitoring Centre). (1992). Global Biodiversity: Status of the Earth's living resources. London: Chapman & Hall. xx + 594 pp.

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