Sewardiella tuberifera Kash. (1915:5)



Taxonomy & Nomenclature



Conservation Status

Rediscovered? (see below)

IUCN RedList status: Vulnerable (2000)


Not recorded since 1984 until it was rediscovered in the Nainital district of Uttarakhand in late September 2021 according to (Pant et al., 2023). However, the species was reported by Pressel et al. (2013) in October 2012 from near Himachal Pradesh's capital Shimla (=Simla) close to the type locality, including a photograph, without any comment on its conservation status. But Pressel et al.'s (2013) report of the species was made in a British magazine which may not have been known to (Pant et al., 2023), so it is unclear whether the 2012 report of the species, nine years earlier than that of (Pant et al., 2023), constitutes the global rediscovery of the species, or whether it is simply a rarely collected species. Given that Pressel et al. (2013) didn't mention it's conservation status, and the large area of which it has been reported, I am inclined to the latter view.



Himachal Pradesh & Uttarakhand, Himalayan India


Anatomy & Morphology



Biology & Ecology










Original scientific description:

Kashyap, S. R. (1915). Morphological and biological notes on new and litle-known West-Himalayan Liverworts. III. New Phytologist 14: 1-18.


Other references:

Chalaud, G. (1932). Mycorhizes et tuberization chez Sewardiella tuberifera Kash. Annales Bryologici 5: 1-16.

Mehra, P. N. (1938). A study of the chromosome number in some Indian members of the family Codoniaceae. ProceedingsIndian AcademySciences B 8: 1-7.

Mehra, P. N. (1977). Cytology of some Indian dioecious and monoecious hepatics. Journ. Hattori Bot. Lab. 43: 137-156.

Mehra, P. N. and Khanna, A. L. (1950). Embryology of Sewardiella tuberifera Kash. Botanical Gazette 112(1): 31-42.

Pande, S. K. and Mishra, R. N. (1937). On the morphology of Sewardiella tuberifera Kash. Proceeding 24th Indian Science Congress (Hyderabad): 263.

Pande, S. K., Srivastava, K. P. and Mishra, R. N. (1955). Studies in Indian Metzgerineae-ll. Sewardiella tuberifera Kash. Phytomorphology 5(1): 57-67.

Pant, G. (1983). Threatened bryophytes of Naini Tal, pp. 313-317. In: Jain, S. K. and Rao, R. R. (eds.). An Assessment of Threatened Plants of India (Proceedings of Seminar, sept. 14-17, 1981). Botanical Survey of India, Botanic Gardens, Howrah,334 pp.

Pant, G., Tewari, S. D. and Joshi, S. (1994). Vanishing greenery in Kumaon Himalaya: Observations on bryoflora. Geophytology 23(2): 253-257.

Pant, S, S.D. Tewari, P. Joshi, M. Bhandari & R. Arya (2023). Rediscovery of Sewardiella tuberifera Kash., a long-lost monotypic endemic Indian liverwort. Journal of Threatened Taxa 15(2): 22726-22730. htps://

Pressel, Silvia, Duckett, Jeff and Villarreal, Juan Carlos. (2013). Hornwort Heaven. Field Bryology 110: 39-46.

Singh, D.K. (2008). Red listing of Hepaticae and Anthocerotae in India, pp. 451-458. In: Mohamed, H., B. B. Baki, A. Nasrulhaq- boyce & P.K.Y. Lee (eds.). Bryology in the new Millennium. University of Malaya, Kuala Lumpur, 513 pp.

Söderström, Lars et al. (2016). World checklist of hornworts and liverworts. PhytoKeys 59: 1-828.

Tewari, S. D. and Pant, G. (1984). Present distribution of two rare, monotypic, endemic liverworts, Stephensoniella brevipedunculata Kash. and Sewardiella tuberifera Kash. in Naini Tal and environs. Journal Himalayan Research and Development 3(1): 48-50.

Udar, R. and Srivastava, S. C. (1983a). Rare and endangered liverworts of India, pp. 303-312. In: Jain, S.K. & R.R. Rao (eds.). An Assessment of Threatened Plants of India (Proceedings of Seminar, sept. 14-17, 1981). Botanical Survey of India, Botanic Gardens, Howrah, 334 pp.

Udar, R. and Srivastava, S. C. (1983b). Reproductive biology of some Indian liverworts. Phytomorphology 33: 37-46.


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