Plagiochila columbiana A.Evans (1896:189)



Taxonomy & Nomenclature

"Most species of Plagiochila exhibit considerable variation in dentition of leaves. The description suggests that P. columbiana is very likely a weak form of P. asplenioides with slightly toothed or entire leaves."

Source: NatureServe


Conservation Status

Missing or invalid (synonym of P. asplenoides)

Last record: 5 December 1987?


Not found during searches of all known historical sites in 1994-95 (Hicks & Amoroso, 1996).



North Carolina & Washington, USA (contra Frye & Clark, 1945:442)


Recorded from Pennsylvania and Virginia by (Frye & Clark, 1945:442), but these records are considered doubtful (Schuster, 1980:383; Stotler & Crandall-Stotler, 2017:665).


Biology & Ecology










Original scientific description:

Evans, A. W. (1896). Notes on the North American species of Plagiochila. Botanical Gazette 21(4): 185-194.


Other references:

Frye, T. C. and Clark, L. (1945). Hepaticae of North America. Part III. Univ. Wash. Publ. Biol. 6: 337-564.

Hicks, M. L. and Amoroso, J. L. (1996). Broyphyte status survey: Plagiochila columbiana Evans. North Carolina Natural Heritage Program and Endangered Species Field Office, US Fish and Wildlife Service, Asheville, North Carolina. Revised 1997. 5 pp. and addenda.

Schuster, R. M. (1980). The Hepaticae and Anthocerotae of North America, East of the Hundredth Meridian, Vol. IV. Columbia University Press, New York.

Söderström, Lars et al. (2016). World checklist of hornworts and liverworts. PhytoKeys 59: 1-828.

Stotler, Raymond E. and Crandall-Stotler, Barbara. (1977). A checklist of liverworts and hornworts of North America. The Bryologist 80(3): 405-428.

Stotler, Raymond E. and Crandall-Stotler, Barbara. (2017). A synopsis of the liverwort flora of North America north of Mexico. Annals of the Missouri Botanical Garden 102(4): 574-709.


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