Plagiochila columbiana A.Evans (1896:189)
Taxonomy & Nomenclature
"Most species of Plagiochila exhibit considerable variation in dentition of leaves. The description suggests that P. columbiana is very likely a weak form of P. asplenioides with slightly toothed or entire leaves."
Source: NatureServe
Conservation Status
Missing if valid (NatureServe, 2026)
Not found during searches of all known historical sites in 1994-95 (Hicks & Amoroso, 1996).
Distribution & Habitat
North Carolina & Washington, USA (contra Frye & Clark, 1945:442)
Recorded from Pennsylvania and Virginia by (Frye & Clark, 1945:442), but these records are considered doubtful (Schuster, 1980:383; Stotler & Crandall-Stotler, 2017:665).
Biology & Ecology
Hypodigm
Media
References
Original scientific description:
Evans, A. W. (1896). Notes on the North American species of Plagiochila. Botanical Gazette 21(4): 185-194.
Other references:
Brinda, John C. and Atwood, John J. (eds.). (2026). The Bryophyte Nomenclator, 14 Jul 2026. Available at: https://www.bryonames.org/ [Accessed on 17 August 2026]
Frye, T. C. and Clark, L. (1945). Hepaticae of North America. Part III. Univ. Wash. Publ. Biol. 6: 337-564.
Hicks, M. L. (1992). Guide to the Liverworts of North Carolina. Duke University, Durham & London. vii + 239 pp.
Hicks, M. L. and Amoroso, J. L. (1996). Broyphyte status survey: Plagiochila columbiana Evans. North Carolina Natural Heritage Program and Endangered Species Field Office, US Fish and Wildlife Service, Asheville, North Carolina. Revised 1997. 5 pp. and addenda.
NatureServe. (2026). Plagiochila columbiana. NatureServe Network Biodiversity Location Data accessed through NatureServe Explorer [web application]. NatureServe, Arlington, Virginia. Available at: https://explorer.natureserve.org/ [Accessed 17 August 2026]
Schuster, R. M. (1980). The Hepaticae and Anthocerotae of North America, East of the Hundredth Meridian, Vol. IV. Columbia University Press, New York.
Söderström, Lars, A. Hagborg, M. von Konrat, S. E. Bartholomew-Began, D. Bell, L. Briscoe, E. A. Brown, D. C. Cargill, D. P. da Costa, B. J. Crandall-Stotler, E. D. Cooper, G. Dauphin, J. J. Engel, K. Feldberg, D. S. Glenny, S. R. Gradstein, X.-L. He, J. Heinrichs, J. Hentschel, A. L. Ilkiu-Borges, T. Katagiri, N. A. Konstantinova, J. Larraín, D. G. Long, M. Nebel, T. Pócs, F. Puche, M. E. Reiner-Drehwald, M. A. M. Renner, A. A. Sass-Gyarmati, A. Schäfer-Verwimp, J. G. Segarra-Moragues, R. E. Stotler, P. Sukkharak, B. M. Thiers, J. Uribe-M., J. Váňa, J. C. Villarreal A., M. J. Wigginton, L. Zhang and R.-L. Zhu. (2016). World checklist of hornworts and liverworts. PhytoKeys 59: 1-828. https://doi.org/10.3897/phytokeys.59.6261
Stotler, Raymond E. and Crandall-Stotler, Barbara. (1977). A checklist of liverworts and hornworts of North America. The Bryologist 80(3): 405-428. https://doi.org/10.2307/3242017
Stotler, Raymond E. and Crandall-Stotler, Barbara. (2017). A synopsis of the liverwort flora of North America north of Mexico. Annals of the Missouri Botanical Garden 102(4): 574-709. https://doi.org/10.3417/2016027
Tropicos. (2026). Tropicos.org. Missouri Botanical Garden. 17 Aug 2026 <https://tropicos.org/name/35185738>
WFO. (2026). Plagiochila columbiana A.Evans. Published on the Internet. Available: https://www.worldfloraonline.org/taxon/wfo-0001196632 [Accessed on 17 August 2026]
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