Monadenia circumcarinata (Stearns, 1879)

Keeled sideband, keeled snail



Taxonomy & Nomenclature



Conservation Status

Rediscovered (Hannah & Smith, 1954; Smith, 1957)


Listed as extinct since 1984 by (Suckling et al., 2004), but this appears to be an error as the species has been found many times since and there is seemingly no report of a second rediscovery of the species after 1954.



California, USA


Biology & Ecology










Original scientific description:

Stearns, R. E. C. (1879). Description of a new species or variety of land snail from California. Annals of the New York Academy of Sciences 1: 395-401.

Other references:

Hannah, G. D. and Smith, A. G. (1954). Rediscovery of two California land snails. Nautilus 67(3): 69-76.

Maciolek, J. A. (1985). Status Survey of the Keeled Snail, Monadenia circumcarinata; Progress report to the Sacramento Endangered Species Office of the USFWS. 9 pp.

Maciolek, J. A. (1986). Status Survey of the Keeled Snail, Monadenia circumcarinata (Stearns 1879). Report to the U.S Fish and Wildlife Service, Region 1, Division of Endangered Species, Sacramento Field Station, Sacramento, CA. 26 pp.

Roth, B. (1972). Rare and endangered land mollusks in California. Sterkiana 48: 1-15.

Smith, A. G. (1957). Snails from California caves. Proceedings of the California Academy of Sciences 29(2): 21-46.

Smith, A. G. (1970). Western land snails. American Malacological Symposium. "Rare and Endangered Mollusks." Malacologia 10(1): 39-46.

Suckling, Kieran, Slack, Rhiwena and Nowicki, Brian. (2004). Extinct and the Endangered Species Act. Centre For Biological Diversity. 63 pp.

Turgeon, D.D., J.F. Quinn, Jr., A.E. Bogan, E.V. Coan, F.G. Hochberg, W.G. Lyons, P.M. Mikkelsen, R.J. Neves, C.F.E. Roper, G. Rosenberg, B. Roth, A. Scheltema, F.G. Thompson, M. Vecchione, and J.D. Williams. (1998). Common and scientific names of aquatic invertebrates from the United States and Canada: Mollusks. 2nd Edition. American Fisheries Society Special Publication 26, Bethesda, Maryland: 526 pp.


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