Pteropus subniger Kerr, 1792:91

Lesser Mascarene flying(-)fox, Small Mauritian flying(-)fox, Dark flying(-)fox, Rougette



Taxonomy & Nomenclature

A list of synonyms can be found here:


Conservation Status


Last record: Mauritius (1859); Réunion (1862); 1864 (Fisher & Blomberg, 2012)

IUCN RedList status: Extinct



Mauritius & Réunion, Mascarene Islands


Biology & Ecology







Above: photograph taken by 'citron'. Released under CC BY-SA 3.0. Available from Wikipedia:



Original scientific description:

Kerr, Robert. (1792). The Animal Kingdom: London: J. Murray & R. Faulder.

Other references:

Bergmans, W. (1990). Taxonomy and biogeography of African fruit bats (Mammalia: Megachiroptera). 3. The genera Scotonycteris Matschie, 1894, Casinycteris Thomas, 1910, Pteropus Brisson, 1762, and Eidolon Rafinesque, 1815. Beaufortia, 40(7):111-177.

Bergmans, W. 1999. Conservation status of African fruit bats (Mammalia, Megachiroptera). In: H. de Iongh and H. Prins (eds), International Seminar on Species Conservation - The IUCN Red List categories discussed, Nederlands Commissie voor Internationale Naturbescherming Medelingen 33, Leiden.

Cheke, A. S. and Dahl, J. F. (1981). The status of bats on Western Indian Ocean islands, with special reference to Pteropus. Mammalia 45(2): 205-238.

Cheke, A. S. and Hume, Julian Pender. (2008). Lost Land of the Dodo: an Ecological History of Mauritius, Réunion & Rodrigues. T & AD Poyser (UK) and Yale University Press (USA).

Fisher, Diana O. and Blomberg, Simon P. (2012). Inferring Extinction of Mammals from Sighting Records, Threats, and Biological Traits. Conservation Biology 26(1): 57-67.

Jones, K.E. et al. 2009 PanTHERIA: A species-level database of life-history, ecology and geography of extant and recently extinct mammals. Ecology 90: 2648.

La Nux, J.B. de, 1772. (Letter on Rousettes and Rougettes). Published, apparently with only a few minor excisions, on pp. 253-262 of Buffon, 1776 (q.v.).

Lee, T. E., Fisher, D. O., Blomberg, S. P. and Wintle, B. A. (2017). Extinct or still out there? Disentangling influences on extinction and rediscovery helps to clarify the fate of species on the edge. Global Change Biology 23(2): 621-634.

Maas, P.H.J. (2011). Lesser Mascarene Flying Fox - Pteropus subniger. In: TSEW (2013). The Sixth Extinction Website. ( Downloaded on 22 December 2013.

Mickleburgh, S., Hutson, A.M., Bergmans, W. & Howell, K. (2008). Pteropus subniger. In: IUCN 2011. IUCN Red List of Threatened Species. Version 2011.1. ( Downloaded on 19 October 2011.

Mickleburgh, S., Hutson, A.M., Bergmans, W. & Howell, K. 2020. Pteropus subniger. The IUCN Red List of Threatened Species 2020: e.T18761A22088168. Accessed on 02 July 2022.

Mickleburgh, Simon P., Hutson, Anthony M. and Racey, Paul A. (1992). Old World fruit bats. An action plan for their conservation. Gland, Switzerland: IUCN. viii + 252 pp.

Simmons, N.B. and A.L. Cirranello. 2023. Bat Species of the World: A taxonomic and geographic database. Version 1.3. Accessed on 06/18/2023.

Turvey, Samuel T. (2009). Holocene mammal extinctions, pp. 41-61. In: Turvey, Samuel T. (ed.). Holocene Extinctions. Oxford, UK & New York, USA: Oxford University Press. xii + 352 pp.

Turvey, Samuel T. and Fritz, Susanne A. (2011). The ghosts of mammals past: biological and geographical patterns of global mammalian extinction across the Holocene. Philosophical Transactions of the Royal Society B 366(1577): 2564-2576. [Supplementary Information]

Van Cakenberghe, V. and Seamark, E. 2012. African Chiropteran Report 2012. African Bats, Pretoria.

Wilson, D.E. & Reeder, D.M. 2005 Mammal species of the world: a taxonomic and geographic reference. Third edition. Baltimore, MD: John Hopkins University Press.



The photograph of the preserved specimen has been released under the [url=]GNU Free Documentation License[/url] by the photographer: citron. Thanks go to the author for their kind permission to place this image in the public domain.


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