Macronycteris besaoka (Samonds, 2007:49)

Big chin horseshoe bat



Taxonomy & Nomenclature

Synonym/s: Hipposideros besaoka Samonds, 2007:49


Conservation Status


Last Record: Late Pleistocene


A recently described species from Late Pleistocene deposits in north-western Madagascar. It's precise age is unknown but certainly not Holocene age when people first colonized Madagascar. Therefore the reason for its extinction remains wide open. But further finds may shed light on its temporal and geographic distribution.



Madagascar (north-western)






Holotype: UA 9478 ("Right maxilla with CP 2,4 M1–3")


For a full list of referred material consult (Samonds, 2007)






Original scientific description:

Samonds, Karen E. (2007). Late Pleistocene bat fossils from Anjohibe Cave, northwestern Madagascar. Acta Chiropterologica 9(1): 39-65.


Other references:

Alumbaugh, Jamie L., Goodman, Steven M. and Samonds, Karen E. (2021). Morphometric Analyses of Modern and Subfossil Macronycteris (Family Hipposideridae) Refine Groups from Anjohibe Cave, Northwestern Madagascar. Acta Chiropterologica 23(1): 33-43. [Abstract]

Goodman, S. M. & Jungers, W. L. Extinct Madagascar: Picturing the island’s past. (University of Chicago Press, 2014).

Goodman, S. M., Ramasindrazana, B., Maminirina, C. P., Schoeman, M. C., & Appleton, B. Morphological, bioacoustical, and genetic variation in Miniopterus bats from eastern Madagascar, with the description of a new species. Zootaxa, 2880(1), 1–19 (2011).

Goodman, Steven M. et al. (2016). How many species of Hipposideros have occurred on Madagascar since the Late Pleistocene? Zoological Journal of the Linnean Society 177(2): 428-449. [Abstract]

Michielsen, Nathan M., Goodman, Steven M., Soarimalala, Voahangy, van der Geer, Alexandra A. E., Dávalos, Liliana M., Saville, Grace I., Upham, Nathan and Valente, Luis. (2023). The macroevolutionary impact of recent and imminent mammal extinctions on Madagascar. Nature Communications 14: 14. [Supplementary Data 1 (mammal list); Supplementary Data 6 (references)]

Patterson, Bruce D. et al. (2020). Evolutionary relationships and population genetics of the Afrotropical leaf-nosed bats (Chiroptera, Hipposideridae). ZooKeys 929: 117-161.

Rakotoarivelo, A. R., Goodman, S. M., Schoeman, M. C. and Willows-Munro, S. (2019). Phylogeography and population genetics of the endemic Malagasy bat, Macronycteris commersoni s.s. (Chiroptera: Hipposideridae).  PeerJ 7: e5866.

Samonds (Irwin), K. E. (2006). The origin and evolution of Malagasy bats: Implications of new Late Pleistocene fossils and cladistic analyses for reconstructing biogeographic history. Ph.D. Dissertation, Department of Anatomical Sciences, Stony Brook University, xx + 403 pp.

Van Cakenberghe, Victor and Seamark, Ernest. (2019). Recent taxonomic changes to african bat taxa (2018-2019). African Bat Conservation News 48: 3-4.


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