Ursus spelaeus ladinicus Rabeder, Hofreiter, Nagel & Withalm, 2004:58



Taxonomy & Nomenclature



Conservation Status


Last record: Late Pleistocene





Biology & Ecology










Original scientific description:

Rabeder, G., Hofreiter, M., Nagel, D., Withalm, G. (2004). New taxa of alpine cave bears (Ursidae, Carnivora). Documents des Laboratoires de géologie Lyon, Hors série, 2: 49-68.


Other references:

Baryshnikov, Gennady F. and Puzachenko, Andrey Yu. (2018). Morphometry of upper cheek teeth of cave bears (Carnivora, Ursidae). Boreas. doi: [Abstract]

Mussi, M. and Palombo, M. R. (2001). Human/carnivore interaction in the Middle Pleistocene of Latium (Central Italy): an open question, pp. 67-75. In: Cavarretta, C., Gioia, P., Mussi, M. and Palombo, M. R. (eds.). The World of Elephants – International Congress, Rome. Proceedings of the 1st international congress – Consiglio nazionale delle ricerche, Rome. [automatic download]

Pérez-Ramos, Alejandro, Romero, Alejandro, Rodriguez, Ernesto and Figueirido, Borja. (2020b). Three-dimensional dental topography and feeding ecology in the extinct cave bear. Biology Letters. doi:

Santos, Elena, Gómez-Olivencia, Asier, Arlegi, Mikel and Arsuaga, Juan Luis. (In Press, 2015). Cranial morphological differences within U. deningeri U. spelaeus lineage: A double traditional and geometric morphometrics approach. Quaternary International. doi:10.1016/j.quaint.2015.11.096 [Abstract]


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