Ursus kanivetz Vereshchagin, 1973



Taxonomy & Nomenclature

Synonym/s: Ursus kanivets Vereshchagin, 1973 [orth. error used by Kosintsev et al., 2022]; Ursus ingressus Rabeder, Hofreiter, Nagel & Withalm, 2004:59


Conservation Status


Last record: Late Pleistocene





Biology & Ecology










Original scientific description:

Vereshchagin, N. K. (1973). Craniometrical characteristic of recent and fossil bears (in Russian). Zoologicheski Zhurnal 52: 920-930.


Other references:

Baca, Mateusz et al. (2016). Retreat and extinction of the Late Pleistocene cave bear (Ursus spelaeus sensu lato). The Science of Nature 103: 92. [Abstract]

Baryshnikov, Gennady F., Gimranov, Dmitry and Kosintsev, Pavel. (In Press, 2018). Variability of the upper incisors in the cave bears (Carnivora, Ursidae) from the Caucasus and Urals. Comptes Rendus Palevol. doi: [Abstract]

Baryshnikov, Gennady F. and Puzachenko, Andrey Yu. (2018). Morphometry of upper cheek teeth of cave bears (Carnivora, Ursidae). Boreas.

Bocherens, H., Stiller, M., Hobson, K. A., Pacher, M., Rabeder, G., Burns, J. A., et al. (2011). Niche partitioning between two sympatric genetically distinct cave bears (Ursus spelaeus and Ursus ingressus) and brown bear (Ursus arctos) from Austria: isotopic evidence from fossil bones. Quaternary International 245(2): 238-248.

Frémondeau, Delphine et al. (In Press, 2020). New mtDNA and Isotopic Evidence on Late Pleistocene Cave Bears in the Balkans: the Case-study of Magura Cave, NW Bulgaria. Acta Zoologica Bulgarica.

Kavcik-Graumann, Nadja et al. (2016). The bears of Illinka Cave near Odessa (Ukraine). ICBS Proceedings, p. 18-25.

Kosintsev, Pavel A. et al. (2022). Mammalian fauna of the Late Pleistocene from the Barsuchiy Dol Cave (Southern Urals). Russian J. Theriol. 21(2): 180-191.

Kosintsev, Pavel A., Simonova, G. V.  and Konovalova, K. Yu. (2023). First Data on Nutrition of the Ural Cave Bear Ursus (Spelaearctos) kanivetz Verestchagin, 1973 (Mammalia, Carnivora, Ursidae) as Based on 13C and 15N Isotope Analyses. Doklady Biological Sciences.

Marciszak, Adrian, Gornig, Wiktoria and Kropczyk, Aleksandra. (2021). Radochowska Cave (Sudety Mts, SW Poland) – one of the European caves with a bear cult?, pp. 62-63. In: Ratajczak-Skrzatek U., Kovalchuk O., Stefaniak K. (eds.). Proceedings of INQUA SEQS 2021 Conference, Wrocław, Poland. University of Wrocław & Polish Geological Society. 119 pp.

Marciszak, Adrian, Gornig, Wiktoria, and Stefaniak, Krzysztof. (2017). Large mammals (carnivores, artiodactyls) from Solna Jama Cave (Bystrzyckie Mts, Southwestern Poland) in the context of faunal changes in the postglacial period of Central Europe. Palaeontologia Electronica 20.1.3A: 1-37.

Marciszak, Adrian Przemysław and Lipecki, Grzegorz. (2020). The history of bears (Ursidae, Carnivora, Mammalia) from Silesia (southern Poland) and the neighbouring areas. Geological Quarterly 64(4): 876-897. doi: 10.7306/gq.1565

Münzel, Susanne C. et al. (2011). Pleistocene bears in the Swabian Jura (Germany): Genetic replacement, ecological displacement, extinctions and survival. Quaternary International 245(2): 225-237.

Münzel, Susanne C. et al. (In Press, 2013). Behavioural ecology of Late Pleistocene bears (Ursus spelaeus, Ursus ingressus): Insight from stable isotopes (C, N, O) and tooth microwear. Quaternary International. [Abstract]

Pedraza, Iván Ramírez et al. (2022). Paleodiet and niche partitioning among the easternmost European cave bears based on tooth wear analysis. Historical Biology 34(6): 1063-1071.

Pérez-Ramos, Alejandro, Romero, Alejandro, Rodriguez, Ernesto and Figueirido, Borja. (2020b). Three-dimensional dental topography and feeding ecology in the extinct cave bear. Biology Letters. doi:

Rabeder, G., Hofreiter, M., Nagel, D., Withalm, G. (2004). New taxa of alpine cave bears (Ursidae, Carnivora). Documents des Laboratoires de géologie Lyon, Hors série, 2: 49-68.

Sivkova, T. N. and Kosintsev, P. A. (2021). First Find of Eggs of the Nematode Baylisascaris transfuga Rudolphi, 1819 (Ascaridoidea, Nematoda) in the Late Pleistocene. Doklady Biological Sciences 499: 103-104.


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