Ursus etruscus Cuvier, 1823

Etruscan bear



Taxonomy & Nomenclature



Conservation Status


Last record: Late Pleistocene





Biology & Ecology










Crégut-Bonnoure, Evelyne and Dimitrijević, Vesna. (2022). New insights on Carnivora (Mammalia) from Trlica (Pljevlja, Montenegro). Revue de Paléobiologie, Genève 41(2): 281-311.

Dong, Wei and Zhang, Yingqi. (2022). Biochronologic sequences of the Pleistocene mammalian faunas in China and correlations with numeric dating. Quaternary Sciences 42(5): 1227-1246.

Gimranov, Dmitry et al. (2022). Ursus etruscus from the late Early Pleistocene of the Taurida сave (Crimean Peninsula). Historical Biology.

Koufos, George D. et al. (2016). Revisiting Ursus etruscus from Greece with description of new material from the Early Pleistocene locality Tsiotra Vryssi (Mygdonia Basin). Closing Symposium, ERC starting grant project "Page" paleoanthropology at the gates of Europe, 1st-4th December, Alte Aula Tübingen.

Koufos, George D. et al. (In Press, 2017). Revisiting Ursus etruscus (Carnivora, Mammalia) from the Early Pleistocene of Greece with description of new material. Quaternary International. [Abstract]

Marciszak, Adrian Przemysław and Lipecki, Grzegorz. (2020). The history of bears (Ursidae, Carnivora, Mammalia) from Silesia (southern Poland) and the neighbouring areas. Geological Quarterly 64(4): 876-897. doi: 10.7306/gq.1565

Mazza, P., Rustioni, M., 1992. Morphometric revision of the Eurasian species Ursus etruscus Cuvier. Palaeontographia Italica 79: 101-146.

Petrucci, Mauro and Sardella, Raffaele. (2009). Ursus etruscus Cuvier, 1823 from the Early Pleistocene of Monte Argentario (Southern Tuscany, Central Italy). Bollettino della Società Paleontologica Italiana 48(2): 89-94.


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