Canis lupus hattai Kishida, 1931:73

Hokkaido wolf, Ezo wolf, Yezo wolf, Yesso wolf, Sakhalin wolf



Taxonomy & Nomenclature

Synonym/s: Canis lupus rex Pocock, 1935; Canis lupus haltai [orth. error used by (Thornback & Jenkins, 1982:307)]


Conservation Status


Last record: 1889 (1887?)

IUCN RedList status: Extinct



Hokkaido, Japan and Sakhalin Island


Biology & Ecology









Original scientific description:

Kishida, K. (1931). Notes on the Yesso wolf. Lansania 3: 72-75.


Other references:

Anonymous. (1970). The Last Wolf of Japan. A Short Story of the Extermination. Selected Catalogue of the Harano Agricultural Museum. Osaka, Japan. [relevant citation?]

Goodwin, Harry A. and Goodwin, J. M. (1973). List of mammals which have become extinct or are possibly extinct since 1600. Int. Union Conserv. Nat. Occas. Pap. 8: 1-20.

Naotaka Ishiguro, Yasuo Inoshima, Nobuo Shigehara, Hideo Ichikawa and Masaru Kato. (2010) Osteological and Genetic Analysis of the Extinct Ezo Wolf (Canis Lupus Hattai) from Hokkaido Island, Japan. Zoological Science 27(4): 320-324.  [Abstract]

Matsubayashi, J., Ohta, T., Takahashi, O. and Tayasu, I. (2017). Reconstruction of the extinct Ezo wolf's diet. Journal of Zoology 302(2): 88-93. [Abstract]

Matsumura, Shuichi, Inoshima, Yasuo and Ishiguro, Naotaka. (2014). Reconstructing the colonization history of lost wolf lineages by the analysis of the mitochondrial genome. Molecular Phylogenetics and Evolution 80: 105-112. [Abstract]

Matsumura, Shuichi, Terai, Yohey, Hongo, Hitomi and Ishiguro, Naotaka. (2020). Analysis of the Mitochondrial Genomes of Japanese Wolf Specimens in the Siebold Collection, Leiden. Zoological Science 38(1). doi: [Abstract]

Mech, L. David. (1974). Canis lupus. Mammalian Species 37: 1-6, 5 figs.

Mech, L. D. and Boitani, L. (IUCN SSC Wolf Specialist Group). (2010). Canis lupus. In: IUCN 2012. IUCN Red List of Threatened Species. Version 2012.2. . Downloaded on 17 March 2013.

Naora, N. (1965). On wolves in Japan. Azekura, Tokyo (in Japanese)

Ohdachi, S. D., Ishiabshi, Y., Iwasa, M. A., Fukui, D. and Saitoh, T. 2015. The Wild Mammals in Japan, Second edition. Shoukadoh, Kyoto, 506 pp.

Sakurai, Ryo et al. (2020). Factors affecting attitudes toward reintroduction of wolves in Japan. Global Ecology and Conservation 22: e01036.

Swancer, Brent. (2009). Japanese Wolves-Part III: The Ezo Wolf. Available online:

Takakura, Tomoaki. (2000). “’Extinct’ wolf may still be on the prowl,” Mainichi Shimbun, November 22. [relevant reference?]

Thornback, Jane and Jenkins, Martin (compilers). (1982). The IUCN Mammal Red Data Book. Part 1: Threatened Mammalian Taxa of the Americas and the Australasian Zoogeographic Region (Excluding Cetacea). Gland, Switzerland: IUCN. 516 pp.

Walker, Brett L. (2004). Meiji modernizatoin, scientific agriculture, and the destruction of Japan’s Hokkaido wolf. Environmental History 9(2): 248-274.

Walker, Brett L. (2008). The Lost Wolves of Japan. University of Washington Press. 360 pp.

Yoneyama, Shoko and Weinstein, Philip. (2024). Reimagining Extinction in Australia and Japan: ‘Voices’ of the Tasmanian Tiger and Hokkaido Wolf. Japanese Studies.


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