Margaritifera marocana (Pallary, 1918)



Taxonomy & Nomenclature

Synonym/s: Margaritifera auricularia marocana (Pallary, 1918)


Conservation Status


IUCN RedList status: Critically Endangered





Biology & Ecology










Original scientific description:

Pallary, P. (1918). Diagnoses d'une cinquantaine de molusques terrestres nouveaux du Nord de l'Afrique. Bulletin de la Société d'Histoire Naturelle de l'Afrique du Nord 9: 137-152.


Other references:

Araujo, R., Toledo, C., Van Damme, D., Ghamizi, M. and Machordom, A. (2009). Margaritifera marocana (Pallary, 1918): a valid species inhabiting Moroccan rivers. Journal of Molluscan Studies 75: 95-101.

Baillie, Jonathan E. M. and Butcher, Ellen R. (2012). Priceless or Worthless? The world’s most threatened species. London, UK: Zoological Society of London.

Fadli M., Ph.D. Thesis, Ibn Tofail University, Kénitra, Morocco, (2003).

Garcia N., Cuttelod A., Abdul Malak D., Gland, Switzerland, Cambridge, UK, and Malaga, Spain. (2010).

Graf, Daniel L. and Cummings, Kevin S. (2021). A ‘big data’ approach to global freshwater mussel diversity (Bivalvia: Unionoida), with an updated checklist of genera and species. Journal of Molluscan Studies 87(1): eyaa034.

Kharboua M., PhD. Marrakech, (1988).

Lopes-Lima, M., Bolotov, I.N., Tu Do, V., Aldridge, D.C., Fonseca, M.M., Ming Gan, H., Gofarov, M.Y., Kondakov, A.V., Prié, V., Sousa, R., Varandas, S., Vikhrev, I.V., Teixeira, A., Wu, R-W., Wu, X., Zieritz, A., Froufe, E., Bogan, A.E. (Accepted, 2018). Expansion and systematics redefinition of the most threatened freshwater mussel family, the Margaritiferidae. Molecular Phylogenetics and Evolution. doi:

Maqboul A., Master’s Thesis, University of Ibn Tofail, Kénitra, Morocco, (1996).

Mohamed bin Zayed Species Conservation Fund. (2015). Annual report 2015. Available from: [Accessed 14 January 2017]

Sousa, R., S. Varandas, A. Teixeira, M. Ghamizi, E. Froufe & M. Lopes-Lima, 2016. Pearl mussels (Margaritifera marocana) in Morocco: Conservation status of the rarest bivalve in African fresh waters. Science of The Total Environment 547: 405-412.

Taybi, A. F., Mabrouki, Y., Ghamizi, M. and Berrahou, A. (2017). The freshwater malacological composition of Moulouya’s watershed and Oriental Morocco. Journal of Materials and Environmental Sciences 8(4): 1401-1416.

Van Damme, D. & Ghamizi, M. 2010. Margaritifera marocana. The IUCN Red List of Threatened Species 2010: e.T184701A8317425. Downloaded on 28 March 2016.


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