Latonia nigriventer Mendelssohn & Steinitz, 1943

Hula painted frog, Israel painted frog, Palestinian painted frog



Taxonomy & Nomenclature

Synonym/s: Discoglossus nigriventer Mendelssohn & Steinitz, 1943


Conservation Status

Last record: 1955

Rediscovered in late 2011

IUCN RedList status: Critically Endangered



Hula Valley, Israel


Biology & Ecology










Original scientific description:

Mendelssohn, H. and Steinitz, H. (1943). A new frog from Palestine. Copeia 1943: 231-233.


Other references:

Andreone, Franco et al. (2022). Threatened and extinct amphibians and reptiles in Italian natural history collections are useful conservation tools. Acta Herpetologica 17(1): 45-58. doi:10.36253/a_h-12349

Biton, R., Boistel, R., Rabinovich, R., Gafny, S., Brumfeld, V. and Bailon, S. (2016). Osteological Observations on the Alytid Anura [i]Latonia nigriventer[/i] with Comments on Functional Morphology, Biogeography, and Evolutionary History. J. Morphol. doi: 10.1002/jmor.20562. [Abstract]

Biton, Rebecca; Geffen, Eli; Vences, Miguel; Cohen, Orly; Bailon, Salvador; Rabinovich, Rivka; Malka, Yoram; Oron, Talya; Boistel, Renaud; Brumfeld, Vlad; Gafny, Sarig. (2013). The rediscovered Hula painted frog is a living fossil. Nature Communications 4. Article 1959. [Abstract]

Day, David. (1981). The Doomsday Book of Animals: A Natural History of Vanished Species. New York, N.Y.: The Viking Press.

IUCN SSC Amphibian Specialist Group. (2012). Discoglossus nigriventer. In: IUCN 2012. IUCN Red List of Threatened Species. Version 2012.2. ( Downloaded on 15 February 2013.

Perl, R. G. Bina et al. (2017). [url=]Natural history and conservation of the rediscovered Hula painted frog, Latonia nigriventer[/url]. Contributions to Zoology, 86(1): 11-37.

Perl, R. G. Bina et al. (2018a). Population genetic analysis of the recently rediscovered Hula painted frog (Latonia nigriventer) reveals high genetic diversity and low inbreeding. Scientific Reports 8: 5588.

Perl, R. G. Bina et al. (2018b). Notes on post-metamorphic colour pattern changes in the Hula painted frog (Latonia nigriventer): How realistic is re-identification of juveniles? Herpetology Notes 11: 475-480. [automatic download]

Regalado, Pedro Galán. (2015). Los Anfibios y Reptiles Extinguidos: Herpetofauna Desaparecida Desde el Año 1500. Monografías de la Universidade da Coruña 155: 1-509.

Renan, Sharon et al. (2017). Living quarters of a living fossil - Uncovering the current distribution pattern of the rediscovered Hula painted frog (Latonia nigriventer) using environmental DNA. Molecular Ecology. DOI: 10.1111/mec.14420

Rinat, Z. 2011. Long thought extinct, Hula painted frog found once again in Israeli nature reserve. Available at: [Accessed 15 February 2013).

Steinitz, H. (1955). Occurrence of Discoglossus nigriventer in Israel. Bulletin of the Research Council of Israel 5B(2)((B)5): 192-193.

Tron, F. (2005). Second Discoglossus nigriventer rediscovery expedition in the central Bekaa Valley, Lebanon: 17-28 April 2005 Expedition Report. A Rocha.


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