Pseudophoxinus handlirschi (Pietschmann, 1933)
Ciçek, Kavinne, Handlirsch's minnow, Eğirdir minnow (shared with P. egirdiri)
Taxonomy & Nomenclature
Synonym/s: Acanthorutilus handlirschi Pietschmann, 1933 (basionym); Oreoleuciscus handlirschi (Pietschmann, 1933); Phoxinellus handlirschi (Pietschmann, 1933)
Conservation Status
Extinct (Küçük, 2012; Freyhof, 2014a,b; Freyhof et al., 2025; Freyhof & Yoğurtçuoğlu, 2026)
Last record: early 1970s or before (Küçük, 2012); 1980's (Freyhof, 2014a,b; Freyhof & Yoğurtçuoğlu, 2026)
IUCN RedList status: Extinct
Distribution & Habitat
Lake Eğirdir, central Anatolia, Turkey
Biology & Ecology
Hypodigm
Media
References
Original scientific description:
Pietschmann, V. (1933). Drei neue Fisharten aus Kleinasien. Mat. nat. KI. Wien 70: 21-23.
Other references:
Akşıray, F. (1961). Bazı Türkiye göllerine aşılanan sudak (Luciopeca lucioperca) balıkları hakkında. İ. Ü. Fen Fak. Hidrobiyoloji Enst. Yayınları IV: 104-113.
Anonymous. (1986). Beyşehir Gölü’nün Limnolojik Özelliklerinin Araştırılması Projesi Sonuç Raporu, T.O.K.B. Isparta İl Müdürlüğü, Isparta.
Baillie, J. and Groombridge, B. (eds). (1996). 1996 IUCN Red List of Threatened Animals. International Union for Conservation of Nature, Gland, Switzerland and Cambridge, UK.
Balik, S. (1995). Freshwater fish in Anatolia, Turkey. Biological Conservation 72: 213-223.
Bogutskaya, N. G. (1992). A revision of species of the genus Pseudophoxinus (Leuciscinae, Cyprinidae) from Asia Minor. Mitt. Hamburg. Zool. Mus. Inst. 89: 261-290.
Campbell, R. N. B. (1992). Food of an introduced population of pikeperch, Stizostedion lucioperca L. in Lake Egirdir, Turkey. Aquac. Fish. Manag. 23: 71-85.
Crivelli, A. J. (1995). Are fish introductions a threat to endemic freshwater fishes in the northern Mediterranean region? Biological Conservation 72: 311–319.
Freyhof, J. (2014a). Pseudophoxinus handlirschi. The IUCN Red List of Threatened Species. Version 2014.1. (http://www.iucnredlist.org). Downloaded on 07 July 2014.
Freyhof, J. (2014b). Pseudophoxinus handlirschi. The IUCN Red List of Threatened Species 2014: e.T40747A19007061. https://dx.doi.org/10.2305/IUCN.UK.2014-1.RLTS.T40747A19007061.en. Downloaded on 26 June 2021.
Freyhof, J. and Özuluğ, M. (2010). Pseudophoxinus fahrettini, a new species of spring minnow from Central Anatolia (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters 20: 325-332.
Freyhof, J. and Yoğurtçuoğlu, B. (2026). Pseudophoxinus handlirschi. The IUCN Red List of Threatened Species 2026: e.T40747A135096365. Accessed on 21 July 2026.
Freyhof, Jörg, Yoğurtçuoğlu, Baran, Jouladeh-Roudbar, Arash and Kaya, Cüneyt. (2025). Handbook of Freshwater Fishes of West Asia. Berlin & Boston: De Gruyter. https://doi.org/10.1515/9783111677811 [p. 376, 377]
Groombridge, B. (ed.). (1994). 1994 IUCN Red List of Threatened Animals. IUCN, Gland, Switzerland and Cambridge, UK.
Harrison, I. J. and Stiassny, M. L. J. (1999). The Quiet Crisis. A preliminary listing of the freshwater fishes of the world that are Extinct or “Missing in Action", pp. 271-331. In: MacPhee, R. D. E. (ed.). Extinctions in Near Time. New York: Kluwer Academic/Plenum Publishers.
Hrbek, T., Stölting, K. N., Bardakci, F., Küçük, F., Wildekamp, R. H. and Meyer, A. (2004). Plate tectonics and biogeographical patterns of the Pseudophoxinus (Pisces: Cypriniformes) species complex of central Anatolia, Turkey. Molecular Phylogenetics and Evolution 32: 297-308.
IUCN. (1990). IUCN Red List of Threatened Animals. IUCN, Gland, Switzerland and Cambridge, UK.
IUCN Conservation Monitoring Centre. (1988). IUCN Red List of Threatened Animals. IUCN, Gland, Switzerland and Cambridge, UK.
Kosswig, C. and Geldiay, R. (1952). Eğirdir Gölü balıkları. balık ve balıkçılık, İst Üniv. Fen Fak. Hidrobiyoloji Arş. Enst. Yayınları 3(1): 3-14.
Küçük, Fahrettin. (1998). Isparta ili içsularında yayılış gösteren tatlısu balıklarının sistematik ve ekolojik özellikleri üzerine araştırmalar. Isparta’nın Dünü, Bugünü, Yarını Semp., 16-17 Mayıs, Isparta.
Küçük, Fahrettin. (2012). Extinct Endemic Fishes of Turkey: Alburnus akili (Gövce) and Pseudophoxinus handlirschi (Kavinne) (Pisces: Cyprinidae). Turkish Journal of Fisheries and Aquatic Sciences 12: 345-347.
Küçük, Fahrettin and İkiz, R. (2004). Fish fauna of streams discharging to Antalya Bay. E.U. J. Fish. Aquat. Sci., 21: 287-294.
Küçük, Fahrettin, Sari, Hasan M., Demir, Orhan and Gülle, İskender. (2009). Review of the ichthyofaunal changes in Lake Eğirdir between 1915 and 2007. Turk. J. Zool. 33: 277-286.
Numann, W. (1958). Anadolu göllerinde limnolojik ve balıkçılık ilmi bakımından araştırmalar ve bu göllerde yaşayan sazanlar hakkında özel bir etüd. İst. Üniv. Fen Fak. Monografi, İstanbul, 7: 114 pp.
Van Neer, W., Wildekamp, R. H., Küçük, F., Ünlüsayin, M., Waelkens, M. and Paulissen, E. (2000). Results of the 1996 survey of the fish fauna of the Aksu River and some lakes in Southwestern Anatolia, and the implications for trade at Sagalassos, In: Waelkens, M. and Loots, L. (eds.). Sagalassos V. Report on the survey and excavation campaigns of 1996 and 1997, Leuven: 828-842.
Van Neer, W., Wildekamp, R.H., Küçük, F. and Ünlüsayın, M. (2008). The 1997-1999 surveys of the Anatolian fish fauna and their relevance to the interpretation of trade at Sagalassos, In: Degryse, P. and Waelkens, M. (eds.). Geo-and Bio-Archeology at Sagalassos and in its Territory, Leuven: 299-323.
Vardakas, Leonidas, Perdikaris, Costas, Freyhof, Jörg, Zimmerman, Brian, Ford, Matthew, Vlachopoulos, Konstantinos, Koutsikos, Nicholas, Karaouzas, Ioannis, Chamoglou, Maria and Kalogianni, Eleni. (2025). Global Patterns and Drivers of Freshwater Fish Extinctions: Can We Learn From Our Losses? Global Change Biology 31(5): e70244. https://doi.org/10.1111/gcb.70244 [Table S1]
Yeğen, V., Balık, S., Bostan, H., Uysal, R. and Bilçen, E. (2006). Recent status of fish fauna in some lakes and dams in lakes region, In: I. proceedings of the national conference on stocking and reservoir management symposium, Antalya-Turkey: 129-139. (in Turkish).
http://fishbase.org/summary/Pseudophoxinus-handlirschi.html
https://extinctanimals.proboards.com/thread/16222/pseudophoxinus-handlirschi-irdir-minnow